Dendrophorbium chopinii Montesinos, 2017

Montesinos-Tubée, Daniel B., 2017, Dendrophorbium chopinii (Compositae: Senecioneae), a new species from Amazonas Region, Peru, Phytotaxa 313 (2), pp. 210-216 : 211

Dendrophorbium chopinii Montesinos , sp. nov. ( Figs 1 , 2 )

Type: — PERU. Amazonas Region: Chachapoyas Province, Leimebamba District, near La Joya archaeological site located in Cerro La Petaca, 1.5 km north of La Joya town, 6°50’04”S, 77°48’20”W, 3468 m, 15 Aug 2016, D. Montesinos & L. García 5010 (holotype HSP-005587, isotypes HUT, F, B).

Description: —Suffruticose perennial plant, erect or rarely prostrate, 0.7–1.5 m tall. Stems covered with black glandular trichomes (0.1–0.5 mm long, 6–8 cells), which turn brownish-red as time passes, mixed with fine, dense, whitish lanuginose trichomes, trichomes somewhat rugulose and lanuginose, ca. 1–8 mm long and 0.1–0.2 mm wide, turning dark red with age. Basal leaves absent. Cauline leaves alternate, decrease in size up the stems, 10–18 cm long, 2–5 cm wide, subcoriaceous, coriaceous with age, elliptic to lanceolate, slightly conduplicate; upper surface lustrous, bright dark green to yellowish green with maturity, rough to rugose with age, covered with thin lanuginose white hairs mixed with dense glands (0.8–1.2 mm long, 6–10 cells), the latter lemon green-coloured and with a red apex, cuticle composed of translucid glands varying from yellow green to bluish-green with age; lower surface light green turning brownish-grey with age, densely covered with a thin indumentum of whitish grey lanuginose hairs, somewhat arachnoid-manicate (1–3 mm long, 0.05–0.1 mm wide) and barely covered with thin glands (<0.1 mm long) turning brownish with age; margin covered with thin lanuginose hairs and glands, irregularly denticulate, with 25–55 denticles on each side (each denticle 1–2 mm long); apex acute, darkish brown-coloured, glabrous; base rounded to subcordate; central nerve prominent, dark red on both surfaces, densely covered by lanuginose hairs, lateral nerves 16–24 pairs on each side of the lamina; petioles 5–15 mm long, conspicuously winged. Synflorescence in large corymbiform panicles, c. 30–60 cm long, 10–30 cm wide, bearing 40 to 70 capitula, with short bracts about 1 cm long. Pedicels 0.8–4.5 cm long, covered with purple-blackish multicellular glands (0.1–0.3 mm long) mixed with a dense tuft of white hairs (2–4 mm). Capitula radiate, 3–3.5 cm in diam., 1–1.8 cm long; involucre campanulate, 8–10 mm in diam., 6–7 mm long. Involucral bracts (phyllaries) 12–14, 5–6 mm long, 1.5–2.1 mm wide, fimbriate, elliptic-oblong, acute-acuminate, lower surface glabrous except at the base and central nerve, upper surface nearly glabrous except the lanuginose base, narrowly pubescent (rarely glabrous) along margins, apex attenuate (1 mm long) bearing a tuft of dense dark purple-blackish multicellular glands scarcely mixed with white multicellular hairs (usually longer than the glands). Ray florets female, 12–14, 10–18 mm long, 3–4 mm wide, elliptic-oblong, bilobulated (slightly curved towards the tip), bright yellow, pale reddish near the apex; floret 4 mm long, style bifurcated, 2.5 mm long, and covered by minute glands at the tip. Disc florets hermaphrodite, ca. 38–42, 10–12 mm long, dark yellow; tube 6–8 mm long, lobes 5, ca. 1 mm long. Anthers crowned by an oblong, obtuse membranous appendage. Style 4–6 mm long, slender, bifurcated, thickened at the base; style branches papillose at the tip. Achenes slightly compressed, heteromorphic. Ray achenes 2–3 mm long, 0.5–0.7 mm wide, with 4–5 longitudinal ribs, furrows between the ribs scarcely papillose; disc achenes 3–5 mm long, 0.7–1 mm wide, with 11–13 longitudinal ribs, furrows between the ribs scarcely papillose. Pappus scabrous, with acutely tipped decurrent spines; ray floret pappus 7–8 mm long and disc floret pappus 7–10 mm long.

Figures

FIGURE 1. Holotype of Dendrophorbium chopinii (HSP-005587).

FIGURE 2. Photographs of Dendrophorbium chopinii. A. Habit (La Joya, Leimebamba, 3468 m). B. Habit (near La Joya, Leimebamba, 3490 m). C. Detail of the internodes and the upper surface of leaves (near Tajopampa, Leimebamba, 3300 m). D. Detail of the lower surface and margin of leaves (near La Joya, Leimebamba, 3490 m). E. Synflorescence in bloom (near La Joya, Leimebamba, 3490 m). F. Dry synflorescence (near Camino Inca, Tajopampa, Leimebamba, 3210 m).