Cryptoneolatirus bellardii (Hörnes, 1854)
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https://doi.org/ 10.11646/zootaxa.5470.1.1 |
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publication LSID |
lsid:zoobank.org:pub:6A4270C2-D3F9-404F-91E7-4A73F2A99AE4 |
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persistent identifier |
https://treatment.plazi.org/id/36508782-FFD0-DB3C-3288-70930869FF09 |
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Plazi |
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scientific name |
Cryptoneolatirus bellardii (Hörnes, 1854) |
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Cryptoneolatirus bellardii (Hörnes, 1854)
Figs 2G View FIGURE 2 , 4F View FIGURE4 , 7A View FIGURE 7 1 –A View FIGURE 1 2 View FIGURE 2
[ Fasciolaria View in CoL ] Bellardii Hörnes — Hörnes 1848: 19 [nomen nudum].
* Fasciolaria Bellardii Hörn. View in CoL —Hörnes 1854: 300, pl. 33, figs 8a–b.
Fasciolaria Bellardii M. Hoern. —Hoernes & Auinger 1890: 264.
Lathyrus (Neolathyrus) bellardii (M. Hö.) — Boettger 1906: 48.
Fasciolaria (Pleuroploca) bellardii Fuchs View in CoL — Sieber 1937: 139.
F [asciolaria]. (P [leuropolca].) bellardii Hörn. View in CoL — Sieber 1958: 152.
Latirus (Neolatirus) danubicus nov. nom. — Strausz 1960: 350.
Latirus (Neolatirus) danubicus Strausz — Nicorici 1972: 68.
Type material. Holotype: specimen illustrated in Hörnes (1854: pl. 33, fig. 8), SL: 27 mm, MD: 12 mm, Baden ( Austria), the specimen is lost.
Illustrated material. NHMW 1865/0001/0211, SL: 35.5 mm, MD: 14.8 mm, Lăpugiu de Sus ( Romania), Figs 2G View FIGURE 2 , 4F View FIGURE4 , 7A View FIGURE 7 1 –A View FIGURE 1 2 View FIGURE 2 .
Revised description. Medium-sized, moderately broad fusiform shell of up to seven teleoconch whorls with conical spire; apical angle 42°. Protoconch conical of 2.5 convex whorls, diameter 950 μm, height 980 μm. Termination of protoconch with weak, opisthocyrt axial riblets. Whorl profile changing from moderately convex on first two teleoconch whorls to very weakly convex on later whorls. First two teleoconch whorls with three broad spiral cords, separated by narrower interspaces, overriding prominent axial ribs, swollen over intersections. Axial ribs fading on third teleoconch whorl. Spiral sculpture on third teleoconch whorl of three broad spiral cords and two weaker secondaries intercalated in interspaces. Abapically cords broaden and flatten, separated by narrow grooves on fourth to fifth whorls. Spiral sculpture becoming subobsolete on penultimate with faint cords restricted to adapical third of last two whorls. Suture very narrowly canaliculate. Last whorl almost flat-sided in subsutural portion, strongly convex and swollen mid-whorl; base very strongly contracting, with weak primary and secondary spiral cords strengthening abapically; fasciole slightly swollen, with weak spiral cords. Aperture pyriform. Columellar callus closely adherent, not thickened. Columella moderately excavated in adapical half, with low, broad parietal fold and second faint fold coinciding with weakly angled transition to siphonal canal in abapical half. Outer lip with four thin, wide-spaced lirae deep inside aperture. Siphonal canal moderately long, moderately narrow, slightly deflected to the left.
Discussion. This species is characterized by its broad conical spire, which distinguish it from Neolatirus recticauda (Fuchs in Karrer, 1877). In addition, Cryptoneolatirus bellardii (Hörnes, 1854) has three strong spiral cords swollen over intersections with numerous axial ribs on early teleoconch whorls, whereas N. recticauda has fewer drop-shaped axial ribs on early whorls overrun by more numerous weaker cords. In addition, the columellar folds are weaker and less numerous in Cryptoneolatirus bellardii (2 vs. 3).
Strausz (1960) assumed that Fasciolaria bellardii Hörnes, 1856 would become a secondary homonym of Turbinella bellardii Michelotti, 1847 if both species would be placed in Fasciolaria or Latirus . Indeed, Boettger (1906) and Strausz (1960) placed Hörnes’ species in Latirus (Neolatirus) [or Lathyrus (Neolathyrus) ] and Bellardi (1884) placed the Italian species in Latirus . Based on this secondary homonymy, Strausz (1960) introduced Latirus (Neolatirus) danubicus as new name for Hörnes’ species. Fasciolaria bellardii Hörnes, 1856 is placed herein in Cryptoneolatirus and Turbinella bellardii Michelotti, 1847 is placed in Polygona ( Van Dingenen et al. 2017) . Therefore, the name Latirus (Neolatirus) danubicus Strausz 1960 is an unnecessary replacement name.
Paleoenvironment. Unknown; probably middle to outer neritic environments.
Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Baden ( Austria) (Hoernes & Auinger 1890); Șimleu Basin: Tusa ( Romania) ( Nicorici 1972); Făget Basin: Lăpugiu de Sus ( Romania) (Hoernes & Auinger 1890).
Genus Eurolatirus nov. gen.
Type species. Eurolatirus ornatus (d’Orbigny, 1852) ; Early Miocene , France .
Diagnosis. Large, broad to moderately slender fusiform shells characterized by strongly convex spire whorls with low-angled, slightly concave subsutural ramp and broad subsutural collar; sculpture of prominent, broad, densely spaced axial ribs overrun by very prominent spiral cords, persisting throughout ontogeny, fasciole broad forming chink or pseudoumbilicus, aperture with denticles on columella and crenulated outer lip with numerous lirae reaching far into aperture; siphonal canal long, broad.
Description. Large, broad to moderately slender fusiform shells with paucispiral protoconch (only known from type species). Spire whorls strongly convex with broad, shallow, slightly concave subsutural ramp and broad subsutural collar. Sculpture of relatively broad, convex axial ribs separated by narrow interspaces, crossed by prominent, narrow, close-set cords of alternating strength. Axial ribs weakening on subsutural ramp and on base, but spiral sculpture strongly developed over entire surface, not weakening with ontogeny. Last whorl attaining ~60–80% of total height, strongly convex, weakly shouldered. Growth lines prominent, interrupting spiral cords. Base strongly constricted, with prominent, relatively widely spaced primary spiral cords, with one or two weaker secondaries intercalated in interspaces. Fasciole prominent, strongly swollen, with strong growth lines, forming distinct chink or pseudoumbilicus. Aperture moderately wide, pyriform. Columellar callus forming broad rim, sharply delimited from base. Columella moderately excavated bearing row of prominent, elongate denticles. Columella bent at transition to siphonal canal. Anal canal moderately incised, parietal denticle very weak; no anal denticle. Outer lip crenulated with numerous prominent lirae starting at edge of aperture, reaching far into aperture. Siphonal canal broad, long, narrow, straight, or slightly deflected to the left, shallowly notched.
Etymology. Combination of Euro - (referring to the European distribution) and Latirus .
Included species. Streptochetus? dispar Peyrot, 1928 ; Fasciolaria valenciennesii Grateloup, 1845 ; Fusus zahlbruckneri Quenstedt, 1884 .
Stratigraphic and geographic range. northeastern Atlantic: Burdigalian (early Miocene) ( Grateloup 1845) and Serravallian (Middle Miocene) ( Peyrot 1928); Central Paratethys Sea: Badenian (Middle Miocene) (hoc opus); Proto-Mediterranean Sea: Serravallian and Tortonian (Middle and Late Miocene) ( Sacco 1904; Landau et al. 2013).
Paleoenvironment. Coastal marine, shallow inner neritic environments, partly with seagrass (own data, M.H.).
Discussion. Eurolatirus nov. gen. is reminiscent of the extant Fusolatirus Kuroda & Habe, 1971 [type species Peristernia pilsbryi Kuroda & Habe, 1952 ; = Fusolatirus coreanicus (E.A. Smith, 1879), by original designation; present-day, Indo-West Pacific]. Fusolatirus , as revised by Snyder & Bouchet (2006), comprises medium sized deep-water species from the Indo-West Pacific with paucispiral protoconch and “ with long, recurved siphonal canal; imbricated subsutural spiral ridge, adpressed to preceding whorl ” (Snyder & Bouchet 2006: 1). Especially Fusolatirus paetelianus ( Kobelt, 1874) is superficially reminiscent of Eurolatirus ornatus (d’Orbigny, 1852) in general shape, apertural features and morphology of the siphonal canal. However, Fusolatirus differs from Eurolatirus in its swollen and more widely spaced axial ribs and Fusolatirus lacks denticles on the columellar callus. A marked difference are the lirae within the outer lip, which start at the edge of the aperture in Eurolatirus but slightly behind the edge in Fusolatirus . In addition, Eurolatirus differs from Fusolatirus in its much larger size. Eurolatirus occurs in shallow water deposits, suggesting an ecological separation from the deep-water genus Fusolatirus .
Eurolatirus dispar ( Peyrot, 1928) was placed by Peyrot (1928) in Streptochetus Cossmann, 1889 with a question mark. This placement has been adopted for Paratethyan occurrences by Strausz (1966) and many subsequent authors including us ( Landau et al. 2013). Fusus intortus Lamarck, 1803 [type species of Streptochetus , original designation by Cossmann (1889: 179), Eocene, Paris Basin] and other Streptochetus species from the Eocene of France, such as S. incertus (Deshayes, 1835) , S. obliquatus (Deshayes, 1835) , S. segregatus (Deshayes, 1865) and S. squamulosus (Deshayes, 1835) , differ quite distinctly from the Miocene species in their comparatively more slender fusiform shells with widely spaced axial ribs, weaker spiral cords, often smooth columella and smooth outer lip lacking denticles, lirae or marginal crenulations. Therefore, the Miocene Eurolatirus is clearly not congeneric with Streptochetus , and we follow Cadée & Janssen (1994), who restricted Streptochetus to Eocene species. Cadée & Janssen (1994) proposed to place ‘ Streptochetus -like’ Miocene species in Streptodictyon Tembrock, 1961 [type species Fusus subelongatus d’Orbigny, 1852 , Rupelian, North Sea]. Later, Lozouet (2015) synonymized both genera with Latirulus Cossmann, 1889 [type species Fusus subaffinis d’Orbigny, 1850 , Ypresian, Paris Basin]. In any case, Latirulus subelongatus (d’Orbigny, 1852) and Latirulus subaffinis ( d’Orbigny, 1850) are obviously unrelated to Eurolatirus and represent slender fusiform shells with weak axial sculpture and delicate spiral cords (see Lozouet 2015: pl. 10, figs 11–12, 14–24).
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Museum Donaueschingen |
| NHMW |
Naturhistorisches Museum, Wien |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Cryptoneolatirus bellardii (Hörnes, 1854)
| Harzhauser, Mathias, Landau, Bernard M. & Vermeij, Geerat J. 2024 |
Latirus (Neolatirus) danubicus Strausz
| Nicorici, E. 1972: 68 |
Latirus (Neolatirus) danubicus
| Strausz, L. 1960: 350 |
Fasciolaria (Pleuroploca) bellardii
| Sieber, R. 1937: 139 |
Lathyrus (Neolathyrus) bellardii (M. Hö.)
| Boettger, O. 1906: 48 |
Fasciolaria
| Hornes, M. 1848: 19 |