Coveenia ulothrix (de Koninck, 1843)

Mottequin, Bernard & Simon, Eric, 2017, New insights on Tournaisian-Visean (Carboniferous, Mississippian) athyridide, orthotetide, rhynchonellide, and strophomenide brachiopods from southern Belgium, Palaeontologia Electronica 37 (3), pp. 1-45 : 33-34

Coveenia ulothrix (de Koninck, 1843)

Figures 4 , 28

* 1843 Terebratula ulothrix de Koninck , p. 635 (replacement name).

1868 Retzia (Terebratula) ulothrix, de Kon. ; Dewalque, p. 332.

1881 Retzia (Terebratula) ulothrix, de Kon. ; Mourlon, p. 46.

v 1887 Retzia ulothrix de Koninck ; de Koninck, p. 92, pl. 22, figs. 1-4.

? 1958 Retzia ulotrix (sic) de Koninck; Demanet, p. 121.

? 1969 Retzia ulotrix (sic) de Kon.; Mortelmans, p. 33.

v 2000 Coveenia ulothrix (de Koninck, 1843) ; Alvarez and Brunton, p. 822, pl. 1, figs. 1- 10, text-fig. 1-2, 5 (see for a more complete synonymy).

v 2007 Coveenia ulothrix (de Koninck, 1843) ; Alvarez, p. 2769, figs. 1851-1852 (copy of Alavarez and Brunton, 2000, pl. 1, figs. 1-4, 6, text-fig. 2 (pro parte).

Material. Besides the material from the RBINS used by Alvarez and Brunton (2000), 18 additional articulated specimens; ULg.PA: four articulated specimens.

Remarks. Although this species has been redescribed thoroughly by Alvarez and Brunton (2000) on the basis of material from Tournai, three additional specimens are illustrated here ( Figure 28.1-18 ) in order to provide the posterior views missing in Alvarez and Brunton’s (2000) publication, and details of the permesothyrid foramen and the symphytium ( Figure 28.16-17 ).

Occurrence. In Belgium, Coveenia ulothrix is known with certainty only from the Tournai area (Tournai Formation; Figure 4 ). Few specimens belonging to the genus Coveenia (C. sp. indet. in Figure 4 ) have been collected in the contemporaneous Yvoir Formation in the Dinant Synclinorium, but further material is required to reach a better identification. Two other Coveenia species are also recognized in Belgium ( Figure 4 ): C. davidsoni ( de Koninck, 1887) (see below) from the Waulsort Formation and C. buchiana (de Koninck, 1843) from the Visé Formation. The genus Coveenia is known from the Tournaisian–Visean of Western Europe, including British Isles ( Alvarez and Brunton, 2000; Alvarez, 2007) but has also been doubtfully reported in the Tournaisian of Algeria by Mottequin and Legrand-Blain (2010).

Figures

FIGURE 4. Distribution of the athyridides, orthotetides and strophomenides described and/or discussed herein within the essentially carbonate Tournaisian and Visean succession of southern Belgium (see text). Stratigraphy, lithostratigraphy (incompletely represented here for the Hainaut and Visé–Maastricht sedimentation areas; formations and members are in roman and italic letters, respectively), and biostratigraphy are adapted from Poty et al. (2002, 2006, 2014). The lenticular Vignobles Member is not represented here as it is rarely developed (it is comprised between the Vaulx Member of the Tournai Formation and the Antoing Formation). Note that the “Grande Brèche” is an informal stratigraphic unit resulting from the dissolution of evaporitic levels in the Grands Malades Formation; other brecciated horizons occur in the Lives Formation.

FIGURE 28. Coveenia ulothrix (de Koninck, 1843) from the Tournai area, Tournai Formation (Tournaisian). 1-5, 16-18, ULg.PA.2016.12.25/10, articulated specimen in ventral, dorsal, lateral, posterior and anterior views, and detail (SEM) of the foramen and the symphytium, and of circular drill hole developed close to the dorsal anterior margin. 6-10, RBINS a13140, articulated specimen in ventral, dorsal, lateral, posterior, and anterior views, Dutoit quarry, Pont-à- Rieu. 11-15, RBINS a13141, articulated specimen, almost complete, in ventral, dorsal, lateral, posterior and anterior views, Dutoit quarry, Pont-à-Rieu. Scale bar equals 10 mm (1-15), 1 mm (16-18).