Colobostema marlettae, Huerta, Heron, 2013

Huerta, Heron, 2013, New species of the genus Colobostema Enderlein (Diptera: Scatopsidae) from Mexico, Zootaxa 3619 (2), pp. 183-194 : 193

publication ID	https://doi.org/ 10.11646/zootaxa.3619.2.6
publication LSID	lsid:zoobank.org:pub:842987A2-768D-4296-A2CD-9FCA8E8F76F1
DOI	https://doi.org/10.5281/zenodo.6162157
persistent identifier	https://treatment.plazi.org/id/03A5E76A-7141-0475-F1C7-FD0BFA53B8B9
treatment provided by	Plazi
scientific name	Colobostema marlettae
status	sp. nov.

Treatment

Colobostema marlettae View in CoL sp. nov.

( Figs. 1 View FIGURE 1 E, 2E, 3F, 4I, J, 5E, 6E, 7E)

Type material. HOLOTYPE male, labeled: MEXICO, Yucatan, Municipio Tizimin, Localidad El Cuyo, cerca al Rancho San Silvestre, selva baja caducifolia, 19-Mar-1996, Malaise trap, Paz Rodríguez, R. & Navarro Correa, C. cols. Specimen dissected and mounted on microscopic slide, deposited in CAIM.

Diagnosis. Deep posteromedian emargination on sternite 7; tergite 7 with sub-basal constrictions, laterally produced, symmetric, with shallow posterior emargination; vestiture of gonocoxite dense and continuous from the lateral base to the apex; tergite 9 rounded apically.

Description. Male. Head brown. Eyes with interommatidial spicules; antenna with 8 flagellomeres ( Fig. 1 View FIGURE 1 E) pale brown; first flagellomere subquadrangular, flagellomeres 2 to 7 broader than long, distal flagellomere clubshaped; flagellum length, 0.50 mm; palpus ( Fig. 2 View FIGURE 2 E) light brown, ovoid, apically with rounded sensory pit, length, 0.06 mm; labella light brown, short and rounded. Thorax brown; anterior spiracular sclerite with 32–33 setae; anepisternum and anepimeron with scattered setae; no katepisternal setae. Legs brownish; femora uniformly brown, tibiae annulated; all tarsi pale brown. Wing ( Fig. 3 View FIGURE 3 F) length approximately 1.60 mm (apex broken in mounting process), width, 0.80 mm; membrane densely micropilose on entire surface. Anterior veins R1, Rs and R4+5 broadened and thickened, with irregularly arranged rows of short setae on dorsal surface; costal cell narrow; C 0.63 of total wing length; r-m very short; fork of M symmetrical; false vein faint; CuA1 ends beyond level of apex of C; CuA2 markedly sigmoid. Halter brown, with two setae on stem. Abdomen: sternite 7 (Fig. 4J) with broad posteromedian emargination, length, 0.42 mm, width, 0.25 mm; tergite 7 (Fig. 4I) with subbasal constrictions, laterally produced, symmetric, with shallow posterior emargination, length, 0.35 mm, width, 0.22 mm. Genital capsule massive ( Fig. 5 View FIGURE 5 E); gonocoxite short and robust, with pointed apex, vestiture dense and continuous from the lateral base to the apex; length, 0.12 mm, width, 0.08 mm; paramere ( Fig. 6 View FIGURE 6 E) bearing comb of dense subapical and apical setae; penis elongate, sinuous subapically, length, 0.1 mm; tergite 9 ( Fig. 7 View FIGURE 7 E) with apex rounded, bearing a pair of strong obtuse tooth-shaped projections, length, 0.16 mm, width at base, 0.18 mm.

Female. Unknown.

Discussion. Colobostema marlettae is very similar to C. marielae and C. lupitae . It is distinguished by the deep posteromedian emargination on sternite 7 (in the other species have a shallow posterior emargination). This group of species shows similarities in shape of tergite 7, male genitalia and setae of paramere. Haenni (1993) describe a group of closely related species characterised by the presence of a group of several flattened setae on apex of paramere among others characters.

As can be seen by the male genital structures (especially the vestiture of parameres and the shape of tergite 9) all these new species form a group of closely related taxa. C. arizonense Cook , C. bijleveldi Haenni and Rapp and C. variatum Cook belong also in this group that appears to be restricted geographically to Mexico, vicinous parts of Nearctic region and the Belize according to our present knowledge (Haenni, pers. obs.).

Distribution. Mexico (Yucatan).

Ecology. Specimen was collected using Malaise trap, in a tropical dry forest, at elevation of 10 m. GPS Coordinates: 21° 27´9´´ N; 87° 42´26´´ W.

Etymology. This species is named after Marlet Trujillo de la Cruz for her support in many aspects of my life.