Coleotichus (Epicoleotichus) borealis Distant, 1899

Tsai, Jing-Fu & Rédei, Dávid, 2010, Taxonomical notes on Oriental and Pacific jewel bugs (Hemiptera: Heteroptera: Scutelleridae), Zootaxa 2572, pp. 25-47 : 26-27

Coleotichus (Epicoleotichus) borealis Distant, 1899 , distinct species

Figs. 1–8, 28–31

Coleotichus borealis Distant, 1899: 31 . Syntypes (2 specimens, erroneously interpreted as holotype and paratype by Cassis & Vanags 2006: 315): “ Formosa ” [= Taiwan], “Lak-ku-li” [= “Rokkiri” (old Japanese transliteration)] [= Liukuei]; BMNH.

Coleotichus excellens (non Walker, 1867): Cassis & Vanags 2006: 315 , partim. Misidentification.

References. Schouteden 1904a: 6 (catalogue, distribution); Schouteden 1905: 320 (in key), 334 (redescription, diagnostic characters, figures); Bergroth 1908: 139 (catalogue); Kirkaldy 1909: 313 (catalogue, distribution); Matsumura 1913: 111 (redescription, habitus); Esaki 1926: 143 (records); Matsumura 1930: 105 (redescription, habitus); Taeuber 1930: 225, 233 (distribution); Matsumura 1931: 1181 (redescription, habitus); Esaki 1932: 1561 (redescription, habitus); Shiraki 1934: 34 (host plant, economic importance); Hoffmann 1935a: 38 (catalogue, distribution); Black 1968: 573 (records, distribution); Göllner- Scheiding 2006: 190 (catalogue, distribution); Cassis & Vanags 2006: 314 (synonymy).

Specimens examined. TAIWAN. Taichung County: “Koroton” [= Fengyuan], [1]907, leg. H. Sauter (1 Ψ, HNHM). Nantou County: “ChipChip” [= Chichi], II. 1909, leg. H. Sauter (2 ɗɗ, HNHM); “Polisha” [= Puli], XII. [1]908, leg. H. Sauter (1 ɗ, HNHM). Hualien County: “Karenko” [= Hualien], 20. VII–4. VIII. [19]19, leg. T. Okuni, J. Sonan, K. Miyake & M. Yoshino (1Ψ, NTU). Chiayi County: “Kagi” [= Chiayi], [1]907, leg. H. Sauter (2 ΨΨ, HNHM [ Figs. 30, 31 ]); same locality, 6. V. 1924, leg. M. Kato (1 ɗ, TARI); “Taihorinsho” [= Dalin], VIII. 1909, leg. H. Sauter (1 ɗ, HNHM). Tainan City: Anping, X. 1909, leg. H. Sauter (1 ɗ, HNHM); Tainan, II. [1]909, leg. H. Sauter (1 Ψ, HNHM). Kaohsiung County: “Kosempo” [= Chiasien], XI. [1]908, leg. H. Sauter (2 ΨΨ, HNHM). Pingtung County: “Koshun” [= Hengchun], III. [1]909, leg. H. Sauter (1 Ψ, HNHM); “Heito” [= Pingtung], 7. X. 1937, collector unknown (1 Ψ, NMNS); same locality, 16. IX. 1939, collector unknown (1 Ψ, NMNS); same locality, 26. VII. 1940, collector unknown (1 ɗ, NMNS); Sichongxi, 8. VII. 1978, leg. K.S. Chang (1 ɗ [Figs. 1–8], NCHU); Hengchun Township, Sheding, 10. VIII. 2004, leg. G.S. Xu (1 ɗ, NPUST); Sheding Sika Deer Restoration Area, 27. III. 2010, canopy sweeping, leg. L.C. Shih (1 Ψ, NCHU).

Discussion. See below under C. excellens .

FIGURES 1–8. Coleotichus borealis Distant, 1899 , male genitalia. 1, pygophore, dorsal view; 2–4, right paramere, three different aspects; 5, phallus, fully inflated, lateral view; 6, same, incompletely inflated, ventral view (CP-II omitted); 7, same, fully inflated, ventral view (apical halves of CP-II1 omitted); 8, apical halves of CP-II1, ventral view. Scales in mm. Lettering: aed: aedeagus sensu stricto; CP-II1 and CP-II2: branches of lateral conjunctival process; CP-III: ventrolateral conjunctival process; mp: median process of cuplike sclerite.

Figures

FIGURES 28 – 35. Coleotichus borealis Distant, 1899 (Taiwan) (28 – 31) and C. excellens Walker, 1899 sensu lato (Australia: Queensland) (32 – 35): 28, 32, head, dorsal view; 29, 33, head, lateral view; 30, 34, gynatrial complex, dorsal view, tergite VIII fully removed; 31, 35, apical receptacle of spermatheca. Scales in mm. Lettering: fec: fecundation sclerite; mp: median pouch of vagina; pt 8: inner surface of paratergite VIII; rs: ring sclerite.