Cochlostoma martorelli montsiccianus

Gofas, Serge, 2001, The systematics of Pyrenean and Cantabrian Cochlostoma (Gastropoda, Cyclophoroidea) revisited, Journal of Natural History 35 (9), pp. 1277-1369 : 1303

Cochlostoma martorelli montsiccianus (Bo®ll, 1890)

Specimens originating from the province of Lerida are rather large (®gure 18a±e), dark and with less conspicuous subsutural colour pattern. The name montsiccianus is given preference over rudicosta because Haas (1929: 398), acting as ®rst revisor, used it as the valid name of a subspecies. In the OrganÄa canyon, there are two coexisting morphs which Fagot (1905) had named:

one resembling the surrounding populations (Boitxols, Jou, OdeÂn...), with dense ribs on which certain white sections form patterns (®gure 18a,b; this is Pomatias organyacus Fagot, 1905 );

one of similar size and shape, but quite smooth and with a uniform reddish brown colour except for whitish remnant ribs along the suture (®gure 18c,d; this is P. alopioides Fagot, 1905 ).

Haas (1929) already questioned whether this variability should deserve taxonomic recognition; sorted lots in museum collections (e.g. Museo de Zoologia , Barcelona) may seem diOEerent but many specimens in the ®eld are transitional. The allozyme data show no basis for separating the two forms: the population is monomorphic for all the loci examined, except LAP for which two alleles are present. LAP B is rare (one B±C heterozygote and 30 CC homozygotes among 31 specimens studied) .