Cochlostoma (T.) hallgassi, Zallot & Kamchev & Schilthuizen & Fehér & Mattia & Gittenberger, 2024

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 104-107

Cochlostoma (T.) hallgassi sp. nov.

urn:lsid:zoobank.org:act:37863EA3-D020-49FF-B4D6-640C063A3B9E

Figs 73I , 82 (blue dot), 92B, 93–94

Differential diagnosis

Cochlostoma (T.) cassiniacum and NFS064 have a different female genital morphology ( Fig. 92D–E ). Cochlostoma (T.) crosseanum ( Paulucci, 1879) can be distinguished because of the spotless shell with widely spaced ribs ( Fig. 92C ), whereas there are well-marked spots and stronger and closely spaced ribs in C. (T.) hallgassi sp. nov. It can be distinguished by the ribbing of the shell (ribs irregular in size in hallgassi , of the same size in mariannae ) and the more curved columellar lobe from C. (T.) mariannae Nordsieck, 2011 ( Fig. 92A ).

Etymology

The species is named after Alessandro Hallgass, a malacologist and friend, who collected most of the samples we have from central Italy.

Types

Holotype

ITALY • ♀; 1- Mount Petrella , 1280 m a.s.l.; 41.3166° N, 13.6542° E; Hallgass leg.; 11 Sep. 2011; RMNH. MOL.507272 .

Paratypes

ITALY • 2 ♀♀; same collection data as for holotype; RMNH. MOL.507273 , RMNH. MOL.507274 • 1 ♂; same collection data as for holotype; RMNH. MOL.507275 .

Other specimens

ITALY • 2- Mt Petrella 1010; 41.3187° N, 13.6175° E; 2012; Hallgass leg.; EZ1118 • 3- Formia ; 41.3069° N, 13.6355° E; 2021; S. Cianfanelli and E. Talenti leg., MZUF 65650 .

Type locality

ITALY • 1- Mount Petrella , 1280 m a.s.l.; 41.3166° N, 13.6542° E .

Description

SHELL. Moderately spaced riblets on last part of protoconch. Teleoconch with two lines of reddish spots on whorls and with ribs not very prominent, rounded and irregular in size. Moderately strong lip with columellar lobe only barely curved toward umbilicus partially covered by it.

MEASUREMENTS. 6 ♀♀: whorls=7.5–8, H = 7.3–8.1 mm, H/W=2.45–2.67, roundness =0.15–0.18, ribs incl.= 64–67, apert. incl.= 14–19°, ribs/mm 1 st wh. =7–11, ribs/mm 4 th wh.=9–13, umb. =narrow slit.

FEMALE GENITAL ORGANS. As in C. (T.) nanum .

Remarks

In the 16S analysis (H3 not amplified), C. (T.) hallgassi is a very close relative (p-distance =0.2%) of a sample from Montenegro, in the Appendix as NFS142. However, the shells of the two samples are different, with marked spots and stronger, narrowly spaced, whitish ribs on the whorls in the Monte Petrella and spotless whorls with weaker ribs of the same color as the background in NFS142.

Figures

Fig. 73. Shell and female genitals of Clade B samples with amplified 16S. A. Cochlostoma (T.) nanum (Westerlund, 1879), Tepke, AT (NHMW-109000/AL/00703). B. C. (T.) nanum, Mt Snežnik 1764, SI (WdM-7005). C. C. (T.) nanum, Mt Risnjak, 1400 m a.s.l., HR (WdM-5666). D. C. (T.) mostarense (Wagner, 1906), Mostar, BeH (EZ-1091). E. C. (T.) elegans (Clessin, 1879), Razbojište near Oltari, HR (WdM-6896). F. C. (T.) stossichi (Hirc, 1881), 2 km NNW of Meja, HR (WdM-1240). G. C. (T.) braueri (Wagner, 1897), Klek Mt, HR (EZ-0068). H. NFS142, Škrčko Jezero, MONT (TxEx-du0044). I. C. (T.) hallgassi sp. nov., Monte Petrella 1010, I (EZ-1118).

Fig. 82.Distributionof samplesin Clade B: red= Cochlostoma (T.)braueri (Wagner, 1897);purple=C. (T.) stossichi (Hirc, 1881); blue=C. (T.) hallgassi sp. nov.; yellow =C. (T.) mariannae Nordsieck, 2011; green =C. (T.) dubium (Wagner, 1897); white = C. (T.) mostarense (Wagner, 1906). Black center for the samples with amplified 16S.

Fig. 92. Turritus Westerlund, 1883 inhabiting the central Apennines. A. Cochlostoma (T.) mariannae Nordsieck, 2011, 1- Vado di Sole, I (EZ-1084). B. C. (T.) hallgassi sp. nov., topotypical specimen, 1- Monte Petrella, I (EZ-1075). C. C. (T.) crosseanum (Paulucci, 1879), 3- Via Salaria, I (EZ-0002). D. C. (T.) cassiniacum (Saint-Simon in Paulucci, 1878), 15- Monte Gennaro, I (EZ-0021). E. NFS064, 1- Rio Fuggio, I (EZ-0022).