Cintaromorpha dacatrai, Häckel & Anichtchenko, 2016

Häckel, Martin & Anichtchenko, Alexander, 2016, Cintaromorpha gen. n. and C. dacatrai sp. n.: a new genus and species of Panagaeini from Pakistan (Coleoptera: Carabidae: Panagaeinae), Baltic Journal of Coleopterology 16 (1), pp. 15-19 : 15-19

Cintaromorpha dacatrai sp. n.

( Fig. 1 a-f)

Holotype (male,“ Pakistan - Kagan V., Murree 2000 m c.a., 6. VII. 1986, C. Dacatra ” (cSD).

Description of holotype. Length 17.5 mm, width 6.5 mm. Proportions: pronotum 1.13x wider than long, 1.83x wider than head, elytra 1.51x wider than pronotum.

Color: Body black, glossy, sparsely covered by black setae; terminal parts of mandibles, basal parts of palps, tarsi and antennae lighter, brownish.

Head long, narrow, 1.83x narrower than pronotum, markedly constricted behind eyes; eyes moderately convex; temples well developed, oblique; labrum strongly excavated at anterior margin; clypeus smooth; vertex convex, sparsely punctured; anterior part of frons smooth, fairly convex at middle, laterally depressed, rugate, lateral margin with sulcus rimming it from mandibular base to posterior margin of eyes, anterior side arcuate, weakly sinuate before eyes, moderately impressed, smooth, sharply bordered externally, medially bordered by a longitudinal ridge separating it medially from elongate lateral pits. Neck smooth.

Pronotum slightly transverse (length to width ratio 1.13), roughly hexagonal, widest just behind midlength, with anterior margin distinctly narrower than base; front angles broadly rounded, indistinct; lateral margins weakly sinuate toward base; hind angles almost rectangular, base straight; lateral rims narrow anteriorly, disappear near front angles; surface densely rugate, coarsely and irregularly punctured; sagittal line distinctly impressed, reaching neither front margin nor base; basal pits elongate, deeply impressed, coarsely punctured.

Elytra weakly ovoid, strongly convex, widest just behind midlength, elongate, weakly tuberose in front of apex; disc weakly flattened; humeri developed but rounded; margins weakly rounded near humeri, widening posterolaterally, strongly rounded toward apex; basal rim indistinct, striae deeply impressed and punctured; intervals conHäckel M., Anichtchenko A.

vex, sparsely punctured, interspaces with dis- rugate, finely rugate at middle, densely covered tinct isodiametric microsculpture. by long setae.

Aedeagus evenly curved, apical lamella short, Venter black, metepisternum anteroposteriorly apex rounded ( Figs.1e, f ).

somewhat longer than wide, trapeziform, Etymology. Named after the collector, Mr. punctured, narrower posteriorly. Ventrites with Stefano Dacatra from Milan, Italy.

anterior margins strongly crenulate, laterally

Distribution. Pakistan: Mansehra District of the Khyber Pakhtunkhwa Province.

Key to Palearctic and Oriental genera of Panagaeini Bonelli, 1810

(Based on recent studies of Häckel & Kirschenhofer, 2014 and Fedorenko, 2015)

1 Unicolorous, dark, apterous species, length> 17 mm ........................................................... 2

- Bicolorous, alate or apterous species; if unicolorous then markedly smaller .................... 3

2 Head broad, not constricted behind eyes, temples short, not reaching half of eye lenght. Pronotum cordate, as wide as long, widest before midlength; front margin wider than base; disc flat, separated from margin by a posteriorly widening rim; lateral margins elevated ......................................................................................................... Cintaroa Kasahara, 1989

- Head elongate, markedly constricted behind eyes, temples long, exceeding half of eye diameter. Pronotum roughly hexagonal, weakly transverse, widest behind midlength; front margin narrower than base; disc arched, lateral margins deflected, anteriorly without an apparent rim, posteriorly rim narrow.........................................… Cintaromorpha gen. n.

3 Paraglossae of the same length as ligula; elytra black, each with two maculae ................ 4

- Paraglossae shorter than ligula, fill lateral margins, ligula shorter. Elytra unicolorous (except genus Panagaeus ) .................................................................................................. 8

4 Protarsi same in both sexes except in Craspedophorus elegans group ......................... 5

- Protarsi of males modified .................................................................................................. 7

5 Penultimate protarsomere not excised, or excised not more than half of its length ......…………... ………………………………………….. Craspedophorus Hope, 1838

- Cleft in penultimate protarsomere exceeds half of protarsomere length ……............… 6

6 Ventrites smooth at bases. Penultimate labial palpomere dilated and plurisetose at inner margin. Monobasic ......................................................................... Dischissus Bates, 1873

- Ventrites with a dense row of large punctures along bases. Penultimate labial palpomere subcylindrical and bisetose at inner margin …………….. Adischissus Fedorenko, 2015

7 Protarsi of males slightly wider than in females, but all tarsomeres lack ventral brushes of setae. Small species less than 8 mm long ...................... Microcosmodes Strand, 1936

- First two male protarsomeres expanded. Monobasic ................. Tinoderus Chaudoir, 1879

8 Cleft in penultimate protarsomere exceeds half of tarsomere length ....................................

.............................................................................................. Euschizomerus Chaudoir, 1850

- Penultimate protarsomere not cleft more than other protarsomeres................................. 9

9 Protarsomeres same in both sexes ...................................... Trichisia Motschulsky, 1860 - At least first protarsomeres of males wider than those in females ................................. 10

10 Only first male protarsomere expanded ................................ Peronomerus Schaum, 1854

- First two male protarsomeres expanded. Elytrons black, each with two yellowish-red

maculae ..................................................................................... Panagaeus Latreille, 1802

Figures

Fig. 1 Cintaromorpha dacatrai sp. nov., HT (male): a. dorsal habitus; b. ventral habitus; c. right metepisternum; d. head, ventral view, detail of mentum, glossae and paraglossae; e. aedeagus, lateral view; f. aedeagus, dorsal view to apex