Chaerilidae Pocock, 1893

Prendini, Lorenzo, Ehrenthal, Valentin L. & Loria, Stephanie F., 2021, Systematics Of The Relictual Asian Scorpion Family Pseudochactidae Gromov, 1998, With A Review Of Cavernicolous, Troglobitic, And Troglomorphic Scorpions, Bulletin of the American Museum of Natural History 2021 (453), pp. 1-153 : 123-124

Family Chaerilidae Pocock, 1893

Chaerilus agilis Pocock, 1899 : MALAYSIA. Habitat: inside cave. Troglomorphies: median and lateral ocelli reduced; pigmentation and sclerotization reduced; pedipalps, legs, and metasoma attenuate. Previous assessments: troglobite (McClure et al., 1967); probably troglophile (Lourenço and Francke, 1985); cavernicoles (Deharveng and Leclerc, 1989); troglophile/trogloxene (Dittmar et al., 2005); suspected of having a subterranean way of life, mainly because found inside caves, but more formal evidence may be necessary (Lourenço and Rossi, 2019). Current assessment: hypogean: troglobite. Citations: Pocock (1899); Bristowe (1952); McClure et al. (1967); Lourenço and Francke (1985); Deharveng and Leclerc (1989); Kovařík (2000b); Dittmar et al. (2005); Lourenço (2008); Beron (2015); Lourenço and Rossi (2019). Remarks: McClure et al. (1967) listed this species as Chaerilus prob. celebensis.

Chaerilus agnellivanniorum Lourenço and Rossi, 2019 : PHILIPPINES. Habitat: inside cave. Troglomorphies: pigmentation and sclerotization reduced; pedipalps, legs, and metasoma attenuate. Previous assessments: may also be a true troglobitic element, but with a less marked degree of regression for several characters (Lourenço and Rossi, 2019). Current assessment: hypogean: troglobite. Citations: Lourenço and Rossi (2019).

Chaerilus cavernicola Pocock, 1894 : INDO- NESIA. Habitat: inside cave. Troglomorphies: pedipalps, legs, and metasoma attenuate. Previous assessments: probably troglophile (Lourenço and Francke, 1985); probably not a troglobite (Volschenk and Prendini, 2008); suspected of having a subterranean way of life, mainly because found inside caves, but more formal evidence may be necessary (Lourenço and Rossi, 2019). Current assessment: hypogean: troglophile (eutroglophile). Citations: Pocock (1894); Wolf (1937); Lourenço and Francke (1985); Kovařík (2000b); Volschenk and Prendini (2008); Lourenço and Rossi (2019).

Chaerilus chapmani Vachon and Lourenço, 1981 : MALAYSIA. Habitat: inside cave. Troglomorphies: median and lateral ocelli reduced; pigmentation and sclerotization absent; pedipalps, legs and metasoma attenuate. Previous assessments: troglobie (Lourenço and Francke, 1985; Lourenço, 1995); cavernicoles (Deharveng and Leclerc, 1989); apparently to some extent troglobitic, a facultative troglobite (Kovařík, 2000a); troglobite (Lourenço, 2008, 2009; Lourenço and Goodman, 2008; Volschenk and Prendini, 2008; Moulds et al., 2013; Lourenço and Rossi, 2019). Current assessment: hypogean: troglobite. Citations: Lourenço and Francke (1985); Deharveng and Leclerc (1989); Lourenço (1995, 2009); Kovařík (2000a, 2000b); Lourenço and Goodman (2008); Volschenk and Prendini (2008); Moulds et al. (2013); Beron (2015); Lourenço and Rossi (2019).

Chaerilus pathom Lourenço and Pham, 2014 : VIETNAM. Habitat: inside cave. Current assessment: hypogean: troglophile (eutroglophile). Citations: Lourenço and Pham (2014b).

Chaerilus sabinae Lourenço, 1995 : INDONE- SIA. Habitat: inside cave. Troglomorphies: median and lateral ocelli absent; pigmentation and sclerotization absent; pedipalps and legs attenuate; telson vesicle enlarged. Previous assessments: cavernicoles, anophtalme (Deharveng and Leclerc, 1989); troglobie (Lourenço, 1995); troglobite (Volschenk and Prendini, 2008; Lourenço, 2009; Lourenço and Rossi, 2019). Current assessment: hypogean: troglobite. Citations: Deharveng and Leclerc (1989); Lourenço (1995, 2009); Kovařík (2000a, 2000b); Volschenk and Prendini (2008); Deharveng and Bedos (2018); Rossi (2018b); Lourenço and Rossi (2019). Remarks: A monotypic genus of dubious validity, Chaerilourencous Rossi, 2018 , was erected to accommodate this species; it is retained in the genus Chaerilus Simon, 1877 in the present contribution.

Chaerilus spinatus Lourenço and Duhem, 2010 : INDONESIA. Habitat: inside cave. Previous assessments: no characteristics of a troglobitic species (Lourenço and Duhem, 2010b); definitely epigean (Lourenço, 2012b). Current assessment: hypogean: troglophile (eutroglophile). Citations: Lourenço and Duhem (2010b); Lourenço (2012b).

Chaerilus telnovi Lourenço, 2009 : INDONE- SIA. Habitat: leaf litter. Troglomorphies: median ocelli absent; lateral ocelli reduced; pigmentation and sclerotization absent; pedipalps, legs, and metasoma attenuate. Previous assessments: litter dwelling (Lourenço, 2009); eyeless forest litter scorpion (Lourenço and Duhem, 2010b); not an epigean species, but a soil dweller (Lourenço, 2012b). Current assessment: endogean. Citations: Lourenço (2009, 2012b); Lourenço and Duhem (2010b); Loria and Prendini (2014); Deharveng and Bedos (2018).