Ceroctis angolensis (Gemminger & Harold, 1870)

Bologna, Marco A., Amore, Valentina & Pitzalis, Monica, 2018, Meloidae of Namibia (Coleoptera): taxonomy and faunistics with biogeographic and ecological notes, Zootaxa 4373 (1), pp. 1-141 : 47-49

Ceroctis angolensis species group

All species belonging to this group are easily distinguishable by the elytral longitudinally vittate pattern and by male genitalia with one single aedeagal distal hook. This group includes at least the following eastern and southern African species: C. angolensis (Gemminger & Harold, 1870) ; C. callicera (Gerstäcker, 1870) ; C. delagoensis (Pic, 1909) ; C. exclamationis ( Marseul, 1872) ; C. matetsiensis Kaszab, 1951 ; C. pilosicollis Borchmann, 1940 ; C. pubicollis Borchmann, 1940 ; C. ruficrus (Gerstäcker, 1854) (variously evaluated in the literature); C. trifurca (Gerstäcker, 1854) ; and C. tripunctata Borchmann, 1940 . Possibly also the following species could belong to the same group: C. bilineata ( Marseul, 1879) ; C. bunkeyana (Pic, 1909) ; C. foveithorax (Pic, 1912) ; C. vittata Borchmann, 1910 .

Ceroctis angolensis (Gemminger & Harold, 1870) ( Fig. 3 A )

Mylabris phalerata Erichson, 1843 nomen praeoccupatum Mylabris angolensis Gemminger & Harold, 1870

Distribution. Angola, Botswana, Namibia , and South Africa.

Material examined and literature records. [Kunene] Epupa: Okakatuwo, 17.4000°S 12.7333°E (SMWN); Hippo Pool on Kunene River 20 km W of Ruacana, 17.4092°S 14.2185°E (CB); Okakatuwo, 17.4500°S 12.7000°E (SMWN); Omunwindi, 17.7833°S 12.8500°E (SMWN). Opuwo: Opuwo town, 18.0500°S 13.8333°E (CB); 6 km NE Orumana, 18.2167°S 13.9500°E (SMWN); C35, Kamanjab-Ruacana 46 km S of cross-road for Opuwo, 18.3280°S 14.3035°E (CB); C 35, 110 km NW Kamanjab, 18.9167°S 14.3833°E (CB). Kamanjab: Beulah 256, 19.6000°S 14.9167°E (SMWN); Kamanjab, 19.6333°S 14.8333°E (SMWN); C40, 26 km E Kamanjab, 19.6957°S 15.1046°E (CB); Otjitambi 25, 19.8167°S 15.1667°E (SMWN); Mooihoek 376, 20.1333°S 15.8833°E (SMWN); Delhi 96, 20.3780°S 15.7280°E (SMWN). Outjo: Ike 346, 19.7533°S 16.6244°E (SMWN); road Outjo-Kalkfeld 16–20 km S Outjo, 20.3333°S 16.1500°E (CB). Khorixas: Fransfontein, 20.1833°S 15.0167°E (SMWN); Duineveld 529, 20.7833°S 14.6333°E (SMWN). [Omusati] Outapi: Mahanene Agric. Research Station, 17.4333°S 14.7833°E (SMWN); Mahanene Res. Station, 17.4500°S 14.8000°E (SMWN). Ogongo: Ogongo Agric. College, 17.6667°S 15.2833°E (SMWN); Ogongo Agric. College, 17.6833°S 15.2833°E (SMWN); Ogongo Agric. College, 17.6833°S 15.3000°E (SMWN). Elim: C46, 15 km NW Oshakati, 17.6833°S 15.5167°E (CB). Tsandi: Tsandi, 17.8333°S 14.9000°E (SMWN). Okahao: C35, Kamanjab-Ruacana 25 km NW of Kamanjab, 18.4775°S 14.6977°E (CB); SW of Dorsland, Etosha N.P., 18.7667°S 14.7333°E (SMWN); Northern border, Bitterwater Firebreak, Etosha N.P., 18.9667°S 15.1500°E (SMWN); Duikersdrink, Etosha N.P., 19.0667°S 14.7167°E (SMWN); Rateldraf, Etosha N.P., 19.0833°S 14.5000°E (SMWN); Otjivasandu, Etosha N.P., 19.2500°S 14.5000°E (SMWN); Kaross, Etosha N.P., 19.3833°S 14.5333°E (SMWN). Ruacana: Ekango, 18.6833°S 14.3167°E (SMWN); NW Etosha N.P., 18.7167°S 14.5833°E (SMWN). [Ohangwena] Okongo: Onamihongwa, 17.6000°S 17.0333°E (SMWN); Etudilondjaba, 17.6000°S 17.6000°E (SMWN). [Kavango] Mpungu: Nangera, 17.6167°S 18.1333°E (SMWN). Mukwe: Popa Falls, 18.1167°S 21.0667°E (SMWN); Popa Falls eastern bank, 18.1167°S 21.5833°E (SMWN); West Caprivi Nature Reserve, 18.1500°S 21.2100°E (SMWN). Ndiyona: Kaudom River, Kaudom Game Reserve, 18.4833°S 20.9333°E (SMWN); Kaudom Camp, Kaudom Game Reserve, 18.5000°S 20.7333°E (SMWN); Kaudom Game Reserve, 18.9333°S 20.9500°E (SMWN); Kaudom Game Reserve, 19.0667°S 20.8000°E (SMWN). [Caprivi] Kongola: Singalamwe, 17.6500°S 23.4167°E (SMWN); Kwando R., 17.7833°S 23.3333°E (SMWN); Manywa Valley, 17.8167°S 23.1500°E (SMWN). Linyandi: Mudumu Nat. Res., 18.1333°S 23.4333°E (SMWN); Mudumu Nat. Res., 18.1667°S 23.4167°E (SMWN); Sangwali, 18.2333°S 23.6000°E (SMWN); 9 km SE Sangwali, 18.3000°S 23.6833°E (SMWN). [Oshana] Okatana: D3609, Oshakati- Ongenga 3 km N of C46 jct.,17.7377°S 15.6727°E (CB). Uuvudhiya: Natukanaoka Pan, Etosha N.P., 18.5833°S 15.7500°E (SMWN); N. of Okandeka, Etosha N.P., 18.9500°S 15.8333°E (SMWN); W of Wolfsnes, Etosha N.P., 19.0500°S 15.8667°E (SMWN). [Oshikoto] Oniipa: B1, 20 km SE Ondangwa, 17.9833°S 16.1333°E (CB). Onayena: B1, Ondangwa-Tsumeb 26 km SE of Ondangwa, 18.0402°S 16.2037°E (CB). Engodi: 19 km ESE Omupaunda, 18.3333°S 16.9333°E (SMWN); 30 km W Oshivelo, 18.4870°S 17.0000°E (CB). Omuthiyagwiipundi: Okashana Rossing Agricultural Centre, 18.4167°S 16.6500°E (SMWN); Fisher Pan, Etosha Nat. Park, 18.7833°S 16.9167°E (SMWN); Gabaub, Etosha N. P., 19.3000°S 16.4167°E (SMWN). Guinas: Operet 312. 18.7333°S 17.2000°E (SMWN); Oshivelo 10 km S on B1, 18.7556°S 17.2420°E (CB); C38, 1 km jct. B1, 18.7725°S 17.2491°E (CB); Ulthoek 770, 19.3300°S 17.6600°E (SMWN). Tsumeb: 5 km E Tsumeb, 19.2600°S 17.7500°E (CB). [Otjozondjupa] Grootfontein: C44, 27 km jct. B8, 19.2428°S 18.7814°E (CB); 10 km NO Grootfontein, 19.5093°S 18.2073°E (CP). Tsumkwe: C44, 52 km jct. B8, 19.2428°S 19.0481°E (CB); 2 km W Omatako, 19.2667°S 19.3000°E (SMWN); Bushmanland, 19.3667°S 19.6000°E (SMWN); Dode, 19.4167°S 20.6000°E (SMWN); Tsumkwe, 19.6000°S 20.5000°E (SMWN); Boesmanland, 19.6000°S 20.6833°E (SMWN); Bushmanland, Aha hills, 19.6167°S 20.9667°E (SMWN); 5 km W Makuri, 19.6667°S 20.6667°E (SMWN); 10 km SE Tsumkwe, 19.6833°S 20.6167°E (SMWN); Naye Naye Pan, 19.7721°S 20.4820°E (SMWN); 2 km NE Namtsoha, 19.7833°S 20.4000°E (SMWN); Gautscha Pan, 19.8000°S 20.5833°E (SMWN); Kremetarkop, 19.8667°S 20.9167°E (SMWN); 3 km W Kuru, 19.9333°S 20.6333°E (SMWN). Otavi: Aigamas 471, 19.4667°S 17.2833°E (SMWN); 5 km E Kombat (road B8), 19.7167°S 17.8000°E (CB). Omatako: Waterberg Pl. Park, 20.3167°S 17.3167°E (SMWN); near Otjahevita road to Grootfontein, 20.3400°S 17.5500°E (CB); Waterberg, Wabi lodge, 20.3429°S 17.5321°E (CP); Waterberg Plateau Park, 20.3667°S 17.3500°E (SMWN); Waterberg Plateau Park, 20.3833°S 17.3000°E (SMWN); Waterberg Plateau Park, 20.4000°S 17.2500°E (SMWN); Waterberg Plateau Park, 20.4167°S 17.2500°E (SMWN); Waterberg Plateau Park, River Omatako, 20.4167°S 17.2500°E (CB); Waterberg Plateau Park, 20.4167°S 17.3333°E (SMWN); Waterberg Plateau Park, 20.4833°S 17.2500°E (SMWN); Rodenstein 307, 20.5626°S 17.2418°E (SMWN); Hamakari 285, 20.5833°S 17.3333°E (SMWN); Hamakari S 373, 20.5833°S 17.4000°E (SMWN); Okosongomingo 149, 20.6167°S 17.1333°E (SMWN); Hamakari 285, 20.6167°S 17.3500°E (SMWN); Otjiwarongo distr., 20.6167°S 17.3500°E (SMWN); Waterberg, jct. roads C22- D2512, 20.6333°S 17.1500°E (CB); Hamakari Sud 373, 20.6667°S 17.3833°E (SMWN); Hamakari 285, 20.6667°S 17.3833°E (SMWN); Hamakari sud 373, 20.6667°S 17.4000°E (SMWN); B1, 28 km S Otjiwarongo, 20.6806°S 16.7787°E (CB); Okonjima 128, 20.8650°S 16.6670°E (SMWN); Okahandja Townlands, 21.6667°S 17.3333°E (SMWN). Otjiwarongo: Cleveland 17. 20.4050°S 16.6590°E (SMWN); Otjowarongo Townlands, 20.4500°S 16.6500°E (SMWN); C63, Outjo-Kalkfeld 15 km N of Kalkfeld, 20.7537°S 16.2210°E (CB). Okakarara: Hereroland West, 20.5167°S 17.4833°E (SMWN); Hereroland West, 20.5500°S 17.4667°E (SMWN); Okakarara, 20.5833°S 17.4333°E (SMWN). Okahandja: Okahandja, 21.9833°S 16.9167°E (CB; SMWN); Damaraland (Okahandja), 21.9833°S 16.9167°E ( Péringuey, 1909); Gross Barmen, 22.1000°S 16.7500°E (CP). [Omaheke] Otjombinde: northern Veter. fence on Game Road, 20.5167°S 20.7000°E (SMWN); Botswana-Namibia border, 2 km northem Veter. fence SWA, 20.5167°S 20.9833°E (SMWN); Eiseb River, 20.6500°S 20.0833°E (SMWN); Confluence Eiseb and Otjinoko Rivers, 20.6667°S 20.1500°E (SMWN); M119 10 km E Helena, 21.8362°S 20.28434°E (CB). Steinhausen: C30, 8 km S jct. C29, 21.8207°S 18.3307°E (CB); Oostenwald, 21.8667°S 19.6333°E (SMWN); Alkm ar 512. 21.8700°S 19.8800°E (SMWN); Owingi 246, 21.9000°S 18.8667°E (SMWN); Kororo Noord 185, 21.9667°S 18.7167°E (SMWN); Eava 383, 22.0833°S 18.9333°E (SMWN); Omitara, C29,15 km jct. D1535, 22.2168°S 18.0407°E (CB); Gobabis, C22, 23 km jct. B6, 22.2533°S 18.9966°E (CB); Geiersberg, Omitara 109, 22.3213°S 18.0331°E (SMWN). Kalahari: M119, 10 km S Botswana border, 22.0238°S 19.9767°E (CB); B6, 20 km W Botswana border, 22.3093°S 19.7964°E (CB); Gobabis, C22, 12 km N jct. B6, 22.3613°S 19.003°E (CB). Gobabis: Gobabis, C22, 2 km N jct. B6, 22.4205°S 19.0037°E (CB); C20, Gobabis-Leonardville 10 km N of Aais, 22.9742°S 18.7280°E (CB). Aminius: Leonardville, 23.5000°S 18.8000°E (SMWN). [Erongo] Dâures: Brandberg, Hungorob Valley, 21.1833°S 14.5167°E ( Bologna, 2000a; SMWN). Omaruru: Otjikoko-Sud 61, 21.2833°S 16.3667°E (SMWN); Omaruru, D2329, 21.4333°S 15.9333°E (CB); C33, 3– 9 km S of Omaruru, 21.4948°S 15.9705°E (CB); 20 km SE Omaruru, 21.6250°S 16.1250°E (JP); C33, 30 km jct. B2, 21.6681°S 15.9799°E (CB); C33, 30– 20 km N Karibib, 21.6833°S 15.9667°E (CB). Karibib: D1935, 30 km NNW Usakos, 21.7833°S 15.5000°E (CB); C33, 17 km jct. B2, 21.839°S 15.9233°E (CB); B2, Usakos- Karibib, 16 km W of Karibib, 21.9358°S 15.7056°E (CB); Karibib, D1953, jct. C32, 21.9552°S 15.8493°E (CB); Okakoara 43, 21.9667°S 16.0167°E (SMWN); C32, 54 km S Karibib, Swakop River, 22.39537°S 15.83432°E (CB). Arandis: 6 km N Arandis, 22.3667°S 14.9833°E (SMWN). [Khomas] Windhoek Rural: Windhoek, vic. Heroes Acre, (OSU); Orumbo 198, 22.3833°S 17.7833°E (SMWN); Windhoek distr., 22.3833°S 17.8333°E (SMWN); Brakwater, 20 km N Windhoek, 22.4000°S 17.0667°E (CB); Richthofen 126, 22.4500°S 17.6000°E (SMWN); Excelsior 286, 22.4500°S 17.6333°E (SMWN); Windhoek Airport, 22.5333°S 17.2500°E (CB); Richthofen 126, 22.5667°S 17.7500°E (SMWN); Alt-Seeis 133, 22.6167°S 17.6333°E (SMWN); Arnhem 222, 22.7000°S 18.1167°E (SMWN); Regenstein 32, 22.7178°S 17.0317°E (SMWN); Gocheganas 26, 22.8333°S 17.1833°E (SMWN); C23, 26 km S Dordabis, 23.0575°S 17.9329°E (CB); C23, 38 km S Dordabis, 23.0715°S 18.0434°E (CB); Kos 28, 23.2667°S 16.1333°E (SMWN); D1228, 7 km ENE Rehobothoad, 23.2833°S 17.2333°E (CB). Windhoek East: Windhoek, 22.5700°S 17.0836°E (CB; SMWN). [Hardap] Rehoboth West Urban: Rehoboth town, 23.3167°S 17.0667°E (CB); Rehoboth, 23.3500°S 17.0667°E (SMWN). Rehoboth Rural: B1, km 21, 23.49991°S 17.12837°E (CB); B1, 28.5 km S Rehobothd, 23.5833°S 17.1500°E (CB); Garies Oos 489, 23.9500°S 16.5500°E (SMWN). Mariental Rural: Stampried, 24.3333°S 18.4000°E (SMWN); Haruchas 156, 24.9500°S 18.8500°E (SMWN); Viljoenskroon 507, 25.1667°S 19.9667°E (SMWN). Gibeon: Wereldend 115, 25.1500°S 16.2333°E (SMWN). [Karas] Berseba: Mukorob 14, 25.5000°S 18.1667°E (SMWN). Keetmanshoop Rural: Koes 202, 25.9500°S 19.1167°E (SMWN); Khabus 146, 26.2833°S 18.2333°E (SMWN); Khabus 146, 26.3000°S 18.2167°E (SMWN); Wildheim Ost 384, 26.4833°S 19.5667°E (SMWN); Karas distr., 27.1352°S 19.4941°E (CB). Lüderitz: near Aus, 26.6667°S 16.2667°E (CP); Diamond Area 1, 27.2500°S 15.7500°E (SMWN). Other records: Swakop River (CB); Ovampoland ( Péringuey 1909); Damaraland ( Kaszab 1956) (ab. damarensis: Kaszab 1956 , Holotype); Namib Sand Sea desert (Seeely 2012); Namibia ( Bologna 2000a; Pitzalis et al. 2014).

Remarks. Types of this species were not examined.

In the checklist of Namibian Meloidae ( Bologna 2000a) , as in most literature, this species is named phalerata . Actually this name is preoccupied by that of an Oriental species now referred to the genus Hycleus ( Pan et al. 2014) .

Ceroctis exclamationis ( Marseul, 1872) ( Fig. 3B )

Mylabris (Ceroctis) exclamationis Marseul, 1872 Mylabris (Ceroctis) bivittata Marseul, 1872

Mylabris (Ceroctis) ngamensis Pic, 1924

Distribution. Angola, Botswana, Mozambique, northeastern Namibia (new species record for this country), South Africa, and Zimbabwe.

Material examined. [Caprivi] Katima Mulilo Urban : Caprivi Katima Mulilo, 17.5000°S 24.2667°E (CB). Kongola: Singalamwe , Eastern Caprivi, 17.6500°S 23.4167°E ( SMWN) ; Kwando R., W Caprivi, 17.7833°S 23.3333°E ( SMWN); 26 km W Kongola , western Caprivi, 17.8000°S 23.0833°E ( SMWN) ; Maywa Valley , westerrn Caprivi, 17.8167°S 23.1500°E ( SMWN) . [Kavango] Mukwe: westerrn Caprivi, 18.0333°S 22.3000°E ( SMWN) ; Omega, westerrn Caprivi, 18.0500°S 22.1833°E (SMWN); Popa Falls , Eeastern Bank, westerrn Caprivi, 18.1167°S 21.5833°E ( SMWN) ; Okavango River, Bagani, Popa Falls , 18.1167°S 21.6333°E (CB; CP).[Oshikoto] Ondangwa-Tsumeb, B 1, 45 km NW Oshivelo Andoni plains , 18.4430°S 16.7342°E (CB). [Otjozondjupa] Tsumkwe: 3 km W Kuru , 19.9333°S 20.6333°E (SMWN).

Other records: Namibia ( Pitzalis et al. 2014) .

Remarks. Types of this species and those of their synonyms were examined at MNHN.

Synonymies must be confirmed after the revision of the genus.

Figures

FIGURE 3. A. Ceroctis angolensis; B. Ceroctis exclamationis; C. Ceroctis bohemani; D. Ceroctis aliena; E. Ceroctis peringueyi; F. Ceroctis trifasciata; G. Ceroctis blanda; H. Ceroctis amphibia; I. Ceroctis karroensis; J. Ceroctis korana; K. Ceroctis braunsiana; L. Ceroctis capensis; M. Ceroctis cfr. seabrai; N. Ceroctis spuria; O. Hycleus dentatus; P. Hycleus tripunctatus; Q. Hycleus jucundus; R. Hycleus peringueyi; S. Hycleus surcoufi; T. Hycleus brincki; U. Hycleus congoensis; V. Hycleus lunatus; W. Hycleus ertli; X. Hycleus arlecchinus; Y. Hycleus basibicinctus.