Cancilla suballigata (Bellardi, 1887)

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 60-62

Cancilla suballigata (Bellardi, 1887)

Figs 15G 1 –G 2 , H 1 –H 2 , I 1 –I 2

[ Nebularia ] striatula Brocc. —Hoernes 1880: 125.

Mitra (Nebularia) striatula Brocchi — Hoernes & Auinger 1880: 80, pl. 9, figs 20a–b [non Brocchi, 1814].

* Mitra suballigata Bell. — Bellardi, 1887a: 71, pl. 4, fig. 15.

Mitra praecedens Bell. — Bellardi, 1887a: 75, pl. 4, fig. 17.

Mitra alligata — Mayer-Eymar 1898: 83.

Mitra suballigata var. transversesulcata Sacc. — Sacco, 1904: 82, pl. 18, fig. 41.

M [itra]. (T [iara].) scrobiculata striatula Brocchi — Sieber 1958a: 154.

Mitra suballigata Bellardi, 1887 — Ferrero Mortara et al. 1981: 159, pl. 48, fig. 12.

Mitra praecedens Bellardi, 1887 — Ferrero Mortara et al. 1981: 160, pl. 48, fig. 13.

Cancilla suballigata (Bellardi, 1887) — Cernohorsky 1991: 37.

Cancilla suballigata (Bellardi, 1887) — Davoli 2000: 191, pl. 5, figs 3–6.

Cancilla suballigata (Bellardi, 1887) — Landau et al. 2013: 212, pl. 33, figs 12–13, pl. 80, fig. 8.

non Mitra (Tiara) striatula (Brocchi) 1814 — Bohn-Havas 1973: 1117, pl. 9, fig. 5 [= Fraudiziba sp. ].

Type material. Syntype ( BS.019.01.151) illustrated in Ferrero Mortara et al. (1981, pl. 48, fig. 12), stored in the Museo Regionale di Scienze Naturali , Torino , Stazzano ( Italy), Tortonian (late Miocene) .

Material. NHMW 1847 /0037/0044b, SL: 30.5 mm , MD: 9.6 mm, Vienna / Pötzleinsdorf , illustrated in Hoernes & Auinger (1880, pl. 9, figs 20a–b), figs 15G 1 –G 2 ; NHMW 2020 /0136/0001, SL: 30.7 mm , MD: 10.4 mm, Bad Vöslau ( Austria) , figs 15H 1 –H 2; NHMW 2020 /0134/0001 SL: 36.6 mm , MD: 9.7 mm, Forchtenau ( Austria) , figs 15I 1 –I 2; NHMW 1863 /0015/0622b, SL: 28.8 mm , MD: 9.2 mm, Niederleis ( Austria) ; NHMW 1855 /0045/0632, 2 specimens, Grund ( Austria) ; NHMW 1846 /0037/0107, 2 specimens, Steinebrunn ( Austria) ; NHMW 1853 /0003/0087, 4 specimens, Forchtenau ( Austria) ; NHMW 1853 /0003/0086, 2 specimens, Forchtenau ( Austria) ; NHMW 1863 /0015/0621, 10 specimens, Niederleis ( Austria) ; NHMW 1847 /0037/0045, Mattersburg ( Austria) ; NHMW 1861 /0001/0425, Marz ( Austria) ; NHMW 1865 /0036/0105, Jerutek at Lysice ( Czech Republic) .

Revised description (of Paratethyan material). Medium sized, moderately slender fusiform shell, with moderately high spire, and high last whorl. Protoconch unknown. About eight teleoconch whorls with deeply incised suture. Spire whorls weakly convex, periphery below mid-whorl. Sculpture consisting of five to six broad, convex to flattish spiral cords separated by narrow grooves with delicate axial riblets. Last whorl faintly shouldered, subcylindrical to convex, with constricted base. About 20 spiral cords on last whorl, more prominent on adapical quarter of whorl, base and fasciole, blurred along periphery. Aperture elongate, posteriorly angulated with indistinct posterior sinus. Columellar callus narrow, indistinct. Columella with three weak columellar folds; fourth abapical fold subobsolete; outer lip thin, Siphonal canal moderately long, twisted, with incised siphonal notch.

Shell measurements and ratios. SL = 28.8–36.6 mm, MD: 9.2–10.4 mm, AA = 24–30°, SL/ MD: 3.2, AL/AW: 5.0, AH/S: 2.1.

Discussion. This species was identified by Hoernes & Auinger (1880) as Mitra striatula Brocchi, 1814 [= Cancilla alligata (Defrance in Blainville, 1825)]. The holotype of Cancilla alligata is a slender fusiform shell with a high conical spire, weakly convex spire whorls, slowly contracting base, indistinct fasciole, and spiral cords of subequal width (see Rossi-Ronchetti 1955, fig. 132; Pinna & Spezia 1978, pl. 67, figs 4–4a). Thus, Cancilla alligata is superficially similar to the Miocene Cancilla suballigata (Bellardi, 1887) , but differs in its slender shell and proportionally higher spire.

Note on Voluta striatula Brocchi, 1814 : Voluta striatula Brocchi, 1814 is a primary homonym of Voluta striatula Schröter, 1804 ( Schröter 1804: 37) and a secondary homonym of Mitra striatula Lamarck, 1811 ( Lamarck 1811: 210) . Therefore, Defrance in Blainville (1825: 494) introduced Mitra alligata as new name. Obviously, Potiez & Michaud (1838: 497) were unaware of Defrance’s name and introduced Mitra brocchii as new name for the same species. Mitra striosa , mentioned by Sismonda (1841: 41) and Hörnes (1852b: 103) as replacement names for Voluta striatula Brocchi , is a nomen nudum and refers to a label name in the Museo Regionale di Scienze Naturali, Torino written by Franco Andrea Bonelli (1784–1830). Therefore, the valid name for the Pliocene species is Cancilla alligata (Defrance in Blainville, 1825). None of the Paratethyan occurrences listed as Mitra striatula in the literature is conspecific with the Pliocene species.

Palaeoenvironment. The assemblage from Vienna /Pötzleinsdorf ( Austria) indicates shallow marine inner neritic conditions (own data), which is in agreement with Turkish occurrences described by Landau et al. (2013).

Distribution in Central Paratethys. Badenian (middle Miocene): North-Alpine-Carpathian Foredeep: Grund ( Austria), Jerutek at Lysice; Vienna Basin: Niederleis, Vienna /Pötzleinsdorf ( Austria) ( Hoernes & Auinger 1880); Eisenstadt-Sopron Basin: Forchtenau, Marz, Mattersburg ( Austria) ( Hoernes & Auinger 1880).

Proto-Mediterranean Sea. Serravallian (middle Miocene): Karaman Basin: Lale, Akboðazi, Akpýnar-Pýnarlar Yaylasý, Seyithasan ( Turkey) ( Landau et al. 2013); Tortonian (late Miocene): Po Basin: Stazzano, Montegibbio (Davoli 20002).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 15. A1–A2. Cancilla wagreichi nov. sp., holotype, NHMW 2020/0130/0001, Lăpugiu de Sus (Romania). B1–B2. Cancilla sismondai (Michelotti, 1847), NHMW 1865/0001/0171, Lăpugiu de Sus (Romania). C1–C2. Cancilla nanostriatula nov. sp., NHMW 2020/0136/0002, paratype, Bad Vöslau (Austria). D1–D2. Cancilla nanostriatula nov. sp., NHMW 1846/0037/0100, holotype, Baden (Austria). E1–E2. Cancilla nanostriatula nov. sp., NHMW 1863/0015/0622a, paratype, Niederleis (Austria). F1–F2. Cancilla nanostriatula nov. sp., paratype, Baden (Austria). G1–G2. Cancilla suballigata (Bellardi, 1887), NHMW 1847/0037/0044b, Vienna/Pötzleinsdorf. H1–H2. Cancilla suballigata (Bellardi, 1887), NHMW 2020/0137/0001, Bad Vöslau (Austria). I1–I2. Cancilla suballigata (Bellardi, 1887), NHMW 2020/0134/0001, Forchtenau (Austria).