Calophya clausa, Burckhardt & Basset, 2000

Burckhardt, D. & Basset, Y., 2000, The jumping plant-lice (Hemiptera, Psylloidea) associated with Schinus (Anacardiaceae): systematics, biogeography and host plant relationships, Journal of Natural History 34 (1), pp. 57-155 : 82-83

Calophya clausa sp. n.

(®gures 4F±K, 12, 57, 58)

Description. Adult. Coloration. Head and body dark brown to black, shiny.

Apex of genal processes light. Forewings transparent, veins ochreous to brown, vein C +Sc almost black. Young specimens with more extended yellow or green coloration.

Structure. Anterior portion of vertex covered in setae which are about as long as distance between them; genal processes long, slender, contiguous in the middle. Forewings (®gure 4F) oval, widest in the middle, narrowly irregularly rounded apically; surface spinules, apart from base of cell cu 2, absent. Genitalia as in ®gure 4G±K. Male subgenital plate short; proctiger massive; paramere long, obliquely truncate apically; distal portion of aedeagus long with straight, long shaft and short apical dilatation. Dorsal margin of female proctiger angularly excavate; subgenital plate pointed apically.

Measurements in mm (5, l 6 m). HW 0.52±0.62; AL 0.54±0.71; WL 1.67±2.33; MP 0.17±0.22; PL 0.13±0.17; AA 0.15±0.17; FP 0.33±0.40; VL/GL 1.00±1.33; AL/F1 2.38±3.33; AL/HW 0.96±1.27; WL/HW 3.21±3.88; WL/WW 2.13±2.48; a/b 1.19±1.61; c/d 0.57±0.95; TL/HW 0.73±0.85; MP/HW 0.33±0.38; AB/AA 1.00±1.13; FP/HW 0.55±0.71; FP/SL 0.80±1.48; FP/CL 2.86±3.27; FP/FA 1.94±2.77.

Fifth instar larva (®gure 12). Yellow, dorsal sclerites dark brown, venter ochreous. Anterior margin of head weakly concavely rounded, not cleft in the middle. Antennae straight, without sectasetae. Legs with claws. Anterior margin of humeral lobes ending level with posterior eye margin. Abdominal dorsum with median row of horns. Caudal plate ratio 0.53. Circumanal ring circular. Head and wing bud margins witout setae; abdominal margin with lanceolate setae.

Host plants, gall and biology. Schinus kauselii , S. montanus , S. patagonicus . Nipple gall on the leaves (®gures 57, 58). On one face with ¯attened, conical nipple bearing hairs at the top around the opening pore. On the other face weakly convex. The gall has one chamber comprising the single larva. For adult emergence, the gall widens at the opening pore but does not tear as in other species such as C. mammifex or C. rubra (®gure 58).

D istribution . Argentina (RõÂo Negro), Chile (Regions V±VII and IX).

Material examined. HOLOTYPE, l Chile: VII Region , Province Talca , Gil de Vilches Park , El Enladrillo, Valle El Venado, 1350±1700 m, 14.i.1996, Schinus patagonicus (D. Burckhardt) , dry mounted ( MHNG).

PARATYPES. Argentina: 2, l 8 m, RõÂo Negro, Bariloche, 5± 10.xi.1926 (F. and M. Edwards) . Chile: V Region , 3, l 1 larva ex nipple gall on leaves, Province Los Andes , 25 km W Portillo, 32ss50¾S 70ss08¾W, 1900±2100 m, 1.xii.1993, Schinus montanus (D. Burckhardt) ; 1, l same data but RõÂo Blanco to Portillo , Valle Aconcagua, 2200±2400 m, 23.xi.1992; 4, l 3 m, 4 larvae, same data but La Campana National Park , Sector Granizo, from leaf galls of Schinus montanus (F. Sa Âiz) ; 2, l 1 m, 4 larvae, same data but La Campana National Park , Sector Ocoa, 1995, Schinus montanus ; VII Region, 4, l 8 m, 5 larvae ex nipple galls on leaves, same data as holotype; 1 larva ex nipple gall on leaf, same data but sector Piedras Blancas, 1350± 1600 m, 12.i.1996, Schinus kauselii ; IX Region, 2, l 1 m, 1 larva ex nipple gall on leaves, same data but Province Malleco, Nahuelbuta National Park, 1300 m, 16± 17.xii.1990, Nothofagus antarctica forest, Schinus patagonicus var. crenuloides ; 3 larvae ex nipple galls on leaf, same data but Province CautõÂn, ConguillõÂo National Park, Playa Linda, 1150 m, 19± 20.xii.1990, Nothofagus antarctica forest, Schinus patagonicus ; 4 larvae ex nipple galls on leaves, same data but 1100 m, 30.i.1996. Dry and slide mounted ( BMNH, MHNG, NHMB, UCVA) .

Material not included in type series. Chile: V Region; larvae from nipple galls of leaves of Schinus montanus , Province Los Andes, 14 km Portillo to RõÂo Blanco, Aconcagua Valley, 2100 m, 23.xii.1995, subalpine scrub (D. Burckhardt); larvae from nipple galls of leaves of Schinus montanus , sama data but 19 km Portillo to RõÂo Blanco, 1900 m; larvae from nipple galls of leaves of Schinus montanus , same data but Portillo to RõÂo Blanco, 1800 m, 31.xii.1995, mediterranean scrub; nipple galls on the leaves, Aconcagua Valley, Juncal, 33ssS, 2200 m, 26.i.1903, Schinus montanus (O. Bucktien) ; nipple galls on the leaves, Province Guillota, La Campana National Park, 1300 m, 11.i.1985 (D. Hollis); VI Region, nipple galls on the leaves of Schinus kauselii and patagonicus , Province Cochagua, Bellavista, RõÂo Claro, 900 m, 11.i.1996, degraded deciduous forest; VII Region, nipple galls on leaves, same data but Province Talca, Gil de Vilches Park, sector Piedras Blancas, 1350±1600 m, 12.i.1996, Schinus kauselii and patagonicus ; nipple galls on the leaves of Schinus patagonicus , same data but sendero El Enladrillado, Valle El Venado, 1350±1700 m, 14.i.1996, mixed Nothof agus forest; IX Region, nipple galls on leaves, Province Malleco, Nahuelbuta National Park, 1300 m, 16± 17.xii.1990, Nothofagus antarctica forest, Schinus patagonicus var. crenuloides (D. Burckhardt) ; larvae from nipple galls on the leaves of Schinus patagonicus , same data but Tolhuaca National Park, sector Laguna Verde, 1000±1300 m, 27.i.1996, mixed Nothof agus and Nothof agus / Araucaria forest; nipple galls on leaves, same data but Province CautõÂn, ConguillõÂo National Park, Playa Linda, 1150 m, 19± 20.xii.1990, Nothofagus antarctica forest, Schinus patagonicus ; larvae from nipple galls on the leaves of Schinus patagonicus , same data but 1100 m, 30.i.1996, mixed Nothof agus / Araucaria forest. Dry or alcohol preserved (BMNH, MHNG).