Burmophilopota wintertoni, Feng & Zhuang & Shih & Ren & Wang, 2024

Feng, Qi, Zhuang, Jialiang, Shih, Chungkun, Ren, Dong & Wang, Yongjie, 2024, The oldest fossil record of Philopotinae (Diptera: Acroceridae) from the mid-Cretaceous amber of Northern Myanmar, Zootaxa 5497 (3), pp. 435-443 : 436-438

Burmophilopota wintertoni sp. nov. ( Figures 1–2 )

urn:lsid:zoobank.org:act:FB30CE5B-FCDD-4B69-BEA0-11835FA6BB15

Material. Holotype, CNU-DIP-MA2015116 ( Figure 1A, B ), male, preserved as a complete individual.

Etymology. The species name is a patronym for Dr. Shaun L. Winterton, in honor of his remarkable contributions to the research of Diptera .

Locality and horizon. Hukawng Valley, Kachin State, northern Myanmar. mid-Cretaceous Cenomanian period, 98.79 ± 0.62 Ma.

Diagnosis. As for the genus.

Description. Body length 11.58 mm, wing length 5.79 mm.

Head: narrower than thorax, spherical. Eyes large, occupying most portion of head capsule; eyes bare and holoptic. Antenna located at mesal of eyes ( Figure 1C ); scape partly hidden by eyes ( Figure 1C ); flagellum with long and two-segmented aristalike stylus ( Figure 1D ). Mouthparts much shorter than head; labellum fleshy ( Figure 1E ); maxillary palpus absent.

Thorax: bare; postpronotal lobe enlarged and fused to single sclerite; postsutural scutum strongly arched and distinctively hump-backed; anepisternum large; katepisternum well-developed; subscutellum bare ( Figure 2A, B ).

Wing: slender, slightly longer than length of abdomen; upper calypter present; lower calypter large, oval; costa circumambient; crossvein h present; Sc long, ending beyond mid-position of costal margin; R 1 long, sinuous, much departing from Sc at wing margin; opening of cell r 1 about 2.5 times as wide as cell sc; Rs short; R 2+3 long, ending before wing apex; R 4+5 with long stalk; R 4 slightly sinuous apically, terminating at wing apex; R 5 ending distinctly beyond wing apex; cell d long; crossvein r-m2 absent; M 1 subparallel to R 5, two times as long as length of M 2; M 3 and M 4 coalescent distally into short petiole; CuP and CuA separate to wing margin; alula well-developed ( Figure 2C–F ).

Legs: hind legs slightly longer than fore- and mid-legs; femur, smooth, equal to length of tibia; tibia with dense short hairs; hind tibia thickened apically; spurs of tibia absent; first tarsus elongated; pretarsus falciform, pulvilli piniform, slightly shorter than claws ( Figure 2G, H ).

Abdomen: stout and bare, shorter than length of wing; terminal position not visible.

Figures

FIGURE 1. Burmophilopota wintertoni gen. et sp. nov., holotype male. (A) Habitus, left lateral view, scale bar = 1 mm; (B) Habitus, right lateral view, scale bar = 1 mm; (C) Head, dorsal view, scale bar = 0.5 mm; (D) Antenna, scale bar = 0.1 mm; (E) Mouthpart, lateral view, scale bar = 0.5 mm. Abbreviations: ar styl: aristalike stylus.

FIGURE 2. Burmophilopota wintertoni gen. et sp. nov., holotype male. (A) Thorax, left lateral view; (B) Thorax, right lateral view; (C–E) Left-wing; (F) Drawing of wing; (G) Tarsus; (H) Claws and pulvilli. All scale bars = 0.5 mm. Abbreviations: al: alula; anepst: anepisternum; CuA: anterior branch of cubital vein; cua: anterior cubital cell; CuP: posterior branch of cubital vein; d: discal cell; kepst: katepisternum; l calyp: lower calypter; M: medial vein; m3: third medial cell; R: radius vein; r-m: radial–medial crossvein; pprn lb: postpronotal lobe; Sc: subcostal vein; sct: scutum; sctl: scutellum; u calyp: upper calypter.