Bonjeania webbi, Winterton, Shaun L., 2007

Winterton, Shaun L., 2007, New species of Acraspisoides Hill & Winterton and Bonjeania Irwin & Lyneborg (Diptera: Therevidae: Agapophytinae), with the description of a new genus, Zootaxa 1438, pp. 1-25 : 18-20

Bonjeania webbi sp. nov.

( Figure 7 )

Holotype male, AUSTRALIA: QUEENSLAND: Carnarvon NP, Mt. Moffatt Summit, 1097m, 25°03’35”S, 148°02’38”E, 20.i.1998, S.L. Winterton, J.&A. Skevington (MEI 91438) ( ANIC).

Paratypes, AUSTRALIA: QUEENSLAND: male, same data as holotype, 27.xi.1997, J.H. Skevington, C.L. Lambkin (MEI 91439) ( QDPI); female, Carnarvon NP, Mt. Moffatt Section, Malaise trap near Mt. Mt. Moffatt Section, 19–23.i.1998, S.L. Winterton, J.&A. Skevington (MEI 91441) ( UQIC); female, Carnarvon NP, Mt. Moffatt Section, Mt. Moffatt Rd., dry creek, 25°03’52”S, 148°01’00”E, 2.xii.1997, J.H. Skevington, C.L. Lambkin, S. Evans, Malaise (MEI 91443) ( INHS); male, Carnarvon NP, Mt. Moffatt Section, W Branch Maranoa R. (site 16), 25°08’22”S, 147°50’37”E, 660m amsl, 26.xi.1995, M.E. Irwin, S.D. Gaimari, ex. 9m Fock’s Malaise trap (MEI 31364) ( ANIC). NEW SOUTH WALES: female, Warrumbungle NP, Buckleys Ck. 1.5 km NE Blackman Camp, 14.i.1994, M.E. Irwin, D.K. Yeates, Malaise trap (MEI 26199) ( MEIC); female, Warrumbungle NP, Browns Ck. 2.5 km N Woolshed, 15.i.1994, M.E. Irwin, D.K. Yeates Malaise trap (MEI 26201) ( MEIC); female, [Warrumbungle NP, Buckleys Ck.] 1.5 km N Camp Blackman, 14.i.1994, M.E. Irwin, D.K. Yeates, Malaise trap (MEI 26200) ( MEIC); female, Warrumbungle NP, Buckleys Ck. 1.5 km NE Blackman Camp, 17.xii.1995, M.E. Irwin (MEI 51082) ( ANIC); female, Warrumbungle NP, Browns Ck. 2.5 km N Woolshed, 16.xii.1995, M.E. Irwin, Malaise trap (MEI 35711) ( ANIC).

Diagnosis. Lower frons and face protruding anteriorly; wing costal margin orange infuscate; scutum bright orange with black, medial stripe; abdomen orange with dark stripe on sternites; male genitalia with epandrium much shorter along midline than width; aedeagus and gonocoxal apodemes elongate; ejaculatory apodemes not enlarged; distiphallus broadly recurved and thickened distally, hypandrium narrow, band-like.

Description. Male. Body length: 8.0–9.0 mm.

Head. Frons slightly narrower than ocellar tubercle at narrowest point; antennal base positioned low on frons ( Fig. 7 A); upper frons flat, lower frons and face protruding anteroventrally; frons and face smooth, glossy black; erect, pale setae sparsely distributed on lower frons; grey pubescence on upper frons below ocellar tubercle; narrow strip of silver velutum laterally on frons along margin of eye, two patches on face beside and below antennal base; ocellar tubercle slightly raised, black with sparse, grey pubescence admixed with erect pale and dark setae posteriorly; occiput concave, black, overlain with grey pubescence; single row of relatively short, black, postocular setae; gena black with long, black setae, silver-white pubescence along margin of eye; palp and labellum brown with scattered, dark setae; antenna brown, length shorter than head; dark setae on scape and pedicel, scape glabrous laterally, flagellum with small patches of short, dark setae basally; flagellum wider and longer than scape and pedicel; scape glossy, pedicel and flagellum overlain with orangebrown pubescence.

Thorax. Scutum and scutellum bright orange with black medial stripe; stripe black, overlain with sparse white-grey pubescence, concentrated along lateral margins of stripe; scutum sparsely covered with elongate, erect, pale and dark setae; scutal macrosetae black; dorsal half of pleuron orange, ventral portion and legs glossy black, erect dark setae on proepisternum, anepisternum, katepisternum, katatergite and coxae, thicker patch of strong, dark setae on katatergite; legs with erect, elongate setae on fore and mid-femora; mid femur with 2–3 strong ventral bristles; fore and mid-tibia dark yellow-brown, hind tibia black; tarsi yellow-cream, individual tarsomeres brown distally; wing hyaline, translucent orange-brown infuscate basally along costal margin; venation yellow; haltere yellow-tan; scutal chaetotaxy (pairs): np, 4–6 [rarely 7]; sa, 1; pa, 1; dc, 2–3; sc, 1.

Abdomen. Bright orange with conspicuous black stripe ventrally; tergites 1–3 brown medially; scattered erect, pale setae on all segments, longer laterally; ventral setae dark; terminalia dark.

Male genitalia. Epandrium much shorter than wide, deeply emarginate on both posterior and anterior margins, strong dark setae laterally ( Fig. 7 B); tergite 8 medially emarginate ( Fig. 7 C); gonocoxal process slightly shorter than gonostylus; inner gonocoxal process equal length to gonostylus ( Figs 7 D–E); gonostylus with patch of setae basally, projecting dorsally; gonocoxal apodeme ca. 2x length of gonocoxite; ventral lobe small, angular; medial atrium large, pubescence anteriorly along medial margin of gonocoxite; hypandrium reduced; sternite 8 broadly quadrangular ( Fig. 7 E); aedeagus elongate, distiphallus with broad ventral curve, compressed laterally, broad apically with minute spines ( Figs 7 F–G); lateral ejaculatory apodeme relatively small in size; ejaculatory apodeme narrow, elongate; arms of ventral apodeme of aedeagal sheath relatively narrow and elongate.

Female. Body length: 7.0– 9.5 mm.

Similar to male except: Frons wider than ocellar tubercle at narrowest point, frons with strip of silver velutum along entire frontal eye margin antennal base orange laterally; setae on thorax, legs and abdomen shorter, often darker than in male.

Female genitalia. Spermathecal sac rounded, single lobe, spermathecal sac duct relatively long, without sclerotised band; single membranous, round spermatheca; spermathecal duct joined to spermathecal sac duct proximal to base of spermathecal sac; accessory gland elongate, relatively small.

Etymology. The specific epithet is a patronym in honour of Donald W. Webb, a prolific New World therevid taxonomist.

Comments. Bonjeania webbi sp. nov. is most closely related to B. bapsis sp. nov. and has a similar distribution throughout inland central and southern Queensland to central New South Wales.

Figures

FIGURE 7. Bonjeania webbi sp. nov.: A, male head, lateral; male genitalia: B, epandrium, dorsal view; C, tergite 8, dorsal view; D, gonocoxite, lateral view; E, gonocoxites and sternite 8, ventral view; F, aedeagus, lateral view; G, same, dorsal view. Scale line = 0.5 mm.