Belisana putao, Zhu & Li, 2021

Zhu, Wenhui & Li, Shuqiang, 2021, Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia, Zootaxa 4963 (1), pp. 115-137 : 125-130

Belisana putao sp. nov.

Figs 9–10 , 13I–J , 15E

Type material. Holotype: Male ( IZCAS Ar41910), roadside between Wasadum and Ziradum (27°29’48.78”N, 97°10’15.90”E, elevation 874 m), Putao, Kachin, Myanmar, 07 May 2017, Z. Chen & J. Wu leg.

Etymology. The specific name refers to the type locality and is a noun in apposition.

Diagnosis. This species resembles B. pianma Huber, 2005 (see Huber 2005: 14, figs 5, 6, 56, 88–92, 112–130) with the elongated distal apophyses on the male chelicerae ( Figs. 10B , 15E ) but can be distinguished by the presence of a subdisto-ventral membranous lamella (arrow 3 in Figs 9C , 13I ) on the procursus, a pair of angular apophyses on the bulbal apophyses (arrow in Figs 10A ), the absence of a ventral hump on the palpal femur and a pair of apophyses on the clypeus.

Description. Male (holotype, IZCAS Ar41910): Total length 1.60 (1.66 with clypeus), carapace 0.66 long, 0.66 wide, opisthosoma 1.00 long, 0.53 wide. Leg I: 11.39 (3.00 + 0.17 + 2.97 + 4.09 + 1.16), leg II: 7.65 (2.03 + 0.15 + 1.91 + 2.72 + 0.84), leg III: 4.85 (1.41 + 0.09 + 1.13 + 1.59 + 0.63), leg IV: 8.04 (2.28 + 0.16 + 1.94 + 2.75 + 0.91); tibia I L/d: 50. Distance PME-PME 0.14, diameter PME 0.06, distance PME-ALE 0.09, AME absent. Habitus as in Figs 10C–E . Carapace pale. Ocular area not elevated. Clypeus unmodified. Chelicerae ( Figs. 10B , 15E ) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.39). Sternum pale, approximately as wide as long (0.41). Thoracic furrow absent. Legs pale. Opisthosoma pale, without spots. Palps as in Figs 9A–B ; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with retrolatero-dorsal apophysis (arrow in Fig 9B ); procursus ( Figs 9A–D ) curved (arrow in Fig. 9A ), complex distally, with droplet-shaped subdisto-dorsal sclerite (arrow 1 in Figs 9C , 13I ), small subdisto-prolateral sclerite (arrow 2 in Figs 9C , 13I ), semicircular subdisto-prolateral membranous lamella (arrow 3 in Figs 9C , 13I ), subdisto-ventral membranous lamella (arrow in Figs 9D , 13J ) and with retrolateral membranous flap (arrow f in Figs 9D , 13J ); bulb ( Fig. 10A ) with hooked apophysis (pair of angular apophyses subdistally, arrow in Figs 10A ) and simple embolus. Retrolateral trichobothria of tibia Iat 6% proximally; legs with short vertical setae on metatarsi, without spines and curved setae; tarsus Iwith 15 distinct pseudopodomeres.

Female: Unknown.

Natural history. The species was found on domed webs attached to the underside of leaves.

Distribution. Myanmar (Kachin, type locality only; Fig. 16 ).

Figures

FIGURE 9. Belisana putao sp. nov., holotype male. A–B. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D).

FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (A–B), 0.20 (C–E).

FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).A–B. B. bachma sp. nov.; C–D. B. cucphuong sp. nov.; E–F. B. jaegeri sp. nov.; G–H. B. kachin sp. nov.; I–J. B. putao sp. nov.; K–L. B. tarang sp. nov. f = flap. Scale bars: 0.05.

FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05.

FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov.