Aviologus duquesnei, Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick & Nel, André, 2014

Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick & Nel, André, 2014, New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera), Zootaxa 3878 (5), pp. 462-470 : 467-468

Aviologus duquesnei sp. nov.

( Fig. 4 )

Material studied: Holotype specimen Avion 0 0 0 34 (print and counterprint), coll. Patrick Roques, to be deposited in the Musée Géologique Pierre Vetter, Decazeville, France.

Age and outcrop. Type locality and horizon. Moscovian (Westphalian C/D equivalent to Bolsovian/Asturian), ‘Terril N°7’, Avion, Pas-de-Calais, France.

Etymology. Named after Mr. Hervé Duquesne, who is a well known collector of plants and insects in the Carboniferous of North of France.

Diagnosis. As for the genus.

Description. A nearly complete forewing perfectly preserved with only fragments of mid part missing. Preserved length 22.6 mm, width 6.9 mm; no defined ScA; ScP anteriorly pectinate, ending on RA, with branches simple or forked and more or less sigmoidal; area between costal margin and ScP 0.8 mm wide; base of RP 5.3 mm distal of wing base; ScP and R (+ M + CuA) basally separated; presence of a common stem R + (M + CuA); RA convex, simple; crossveins in area between RA and RP not preserved, if present; RP rather concave; first fork of RP 6.5 mm from its base, RP with seven branches on wing margin; first branch short before its fusion with MA; CuA separated from M in basal third of wing; M dichotomously ramified into neutral MA and concave MP, far, 5.3 mm, from its base; MA distally fused with first branch of RP, simple; MP with two branches near posterior wing margin; CuA fused with CuPaα at base of M; CuA + CuPaα posteriorly pectinate, with its first posterior branch concave; CuPaα short basal of its fusion with CuA; CuPaβ present, simple; a series of short crossveins in area between CuA + CuPaα and CuPaβ; division of CuP into CuPa and CuPb 2.8 mm from wing base; concave CuPb simple; area between CuPa and CuPb narrow; three anal veins ending on posterior wing margin, all simple and more convex than CuP.

Discussion. This forewing corresponds to that of a clade Panorthoptera for the same reasons as above (except for the ScP ending on RA). The veins CuPaβ and CuA + CuPaα not strongly approximate exclude affinities with the Caloneurodea. Aviologus gen. nov. shares with the Orthoptera the veins CuPaβ, CuPb and 1A simple, and MP divided into two small distal branches. Nevertheless Aviologus differs from the Oedischiidae in the basal fork of M far from point of separation between M and CuA. Also its fusion of MA with first posterior branch of RP is quite different from the stem of RP and MA strongly approximate or fused for a short distance of the Oedischiidae . Aviologus differs from the Geraridae in the three characters: RA without well defined branches, MA fused with first posterior branch of RP instead of being fused or approximating the stem of RP, and CuPb simple. Lastly Heterologus duyiwuer and Heterologus langfordorum share with Aviologus a simple CuPaβ, and a long stem of M before its first fork; but both strongly differ from Aviologus in the median and radial veins well separated and MP with numerous branches. Bruaylogus differs from Aviologus in median and RP veins well separated, MP with numerous branches, M with a long stem, plus the presence of only two anal veins ending on posterior wing margin instead of three in Aviologus . Aviologus corresponds to a new genus but its family affinities remains undetermined since a new phylogenetic analysis is not performed, as for Bruaylogus . It is even not sure if they belong to the same or to two separate families.

Figures

FIGURE 4. Aviologus duquesnei gen. nov. et sp. nov., holotype Avion 0 0 0 34 (print and counterprint) stored in MNHN coll. finally deposited in the Musée Géologique Pierre Vetter, Decazeville, France. A, photograph of forewing venation; B, line drawing of forewing venation. Scale bar = 5 mm.