Aurantilaria pyrulaeformis (Hoernes & Auinger, 1890)

Harzhauser, Mathias, Landau, Bernard M. & Vermeij, Geerat J., 2024, The Dolicholatiridae and Fasciolariidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5470 (1), pp. 1-92 : 10-13

Aurantilaria pyrulaeformis (Hoernes & Auinger, 1890)

Figs 5A–C

* Fasciolaria pyrulaeformis nov. form.—Hoernes & Auinger 1890: 265, pl. 32, figs 5–6.

Fasciolaria (Pleuroploca) pyrulaeformis Hoernes, Auinger, 1980 [sic]— Nikolov 1994: 54, pl. 5, figs 1–2.

Fasc [iolaria]. pirulaeformis [sic] Hoern. et Auinger— Cossmann 1901: 40.

Type material. Lectotype (designated by Nikolov 1994: 54): NHMW 1860 /0040/0177, SL: 105.6 mm, MD: 55.4 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1890: pl. 32, fig. 5), Figs 5A 1 –A 2 . Paralectotypes: NHMW 1866 /0040/0301a, SL: 76.0 mm , MD: 34.3 mm, Lăpugiu de Sus ( Romania), Figs 5B 1 –B 2 . NHMW 1857 /0024/0021, SL: 95.4 mm , MD: 47.5 mm, Lăpugiu de Sus ( Romania), Fig. 5C . NHMW 1866 /0040/0301, SL: 55.3 mm , MD: 26.5 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1890: pl. 32, fig. 6) .

Additional material. 1 spec., NHMW 2023 /0367/0001, Baden-Sooss ( Austria) .

Revised description. Very large, broad fusiform shell of six teleoconch whorls with gradate spire; apical angle ~50°. Protoconch unknown. Early teleoconch whorls convex with shallow, concave subsutural ramp. Broad axial ribs forming blunt nodes along periphery, fading on subsutural ramp. Spiral sculpture of several prominent spiral cords, separated by narrower interspaces. Cords prominent on subsutural ramp and between nodes, subobsolete over nodes. Suture moderately incised, with narrow subsutural collar, bearing two spiral cords. Growth lines more prominent on subsutural ramp and strongly raised between spiral cords of subsutural collar. Nodes at periphery split on fourth and fifth teleoconch whorls by intercalation of narrow spiral cord. Last whorl attaining 75% of total height, with broad, shallow, concave subsutural ramp, roundly angled at strong shoulder, convex below, moderately constricted at base; fasciole moderately broad, poorly delimited. Spiral sculpture of about four narrow, widely spaced spirals over subsutural ramp, five spiral rows of large, low nodes below shoulder; interspaces between spiral rows of nodes widening abapically, bearing one or two secondary cords; cords over base prominent, elevated. Aperture moderately wide, pyriform. Columellar callus forming broad thickened rim along siphonal canal, thinner and adherent above. Columella moderately and broadly excavated with three strongly oblique columellar folds deep inside aperture; only two folds visible in aperture of fully-grown specimens; adapical fold weak, abapical fold marking slightly angled transition to siphonal canal. Anal canal narrowly incised. Outer lip thin, distinctly notched at shoulder, with numerous thin lirae extending deep within aperture. Siphonal canal moderately long, narrow, straight or slightly deflected to the left.

Discussion. This species was placed in Aurantilaria by Snyder et al. (2012: 46), which is followed herein. The early teleoconch whorls of Aurantilaria pyrulaeformis (Hoernes & Auinger, 1890) are nearly indistinguishable from those of Aurantilaria tarbelliana ( Grateloup, 1845) , but later whorls differ in their bifid nodes along the periphery. In addition, the spiral rows of nodes on the last whorl distinguish this species from A. tarbelliana , which has a longer siphonal canal. We are not aware of any closely related species in the European Neogene. So far, A. pyrulaeformis is only known from southern basins of the Central Paratethys Sea and was restricted to the early/middle Badenian (Langhian).

Aurantilaria carminamaris ( Maury, 1917) , from the Messinian Cercado Formation in the Dominican Republic, is a Western Atlantic counterpart of the Paratethyan species but differs in the more pointed nodes and the presence of a single row of nodes along the shoulder (see Landau & Vermeij 2014: pl. 4, figs 21–25).

Paleoenvironment. Unknown; probably middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania) (Hoernes & Auinger 1890); Dacian Basin: Bivolare ( Bulgaria) ( Nikolov 1994).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width, S: siphonal canal.

FIGURE 2. Protoconchs of Paratethyan Fasciolariidae. A. Dulaiania pleurotomoides (Hoernes & Auinger 1890), holotype, NHMW 1855/0043/0041. B. Aurantilaria tarbelliana (Grateloup, 1845), NHMW 1865/0001/0210, Lăpugiu de Sus (Romania). C. Polygona vindobonensis (Csepreghy-Meznerics, 1956), NHMW 2023/0350/0005, Bad Vöslau (Austria). D. Eurolatirus ornatus (d’Orbigny, 1852), NHMW 1846/0037/0257, Gainfarn (Austria). E. Tarantinaea hoernesii (Seguenza, 1875), NHMW 1853/0002/0052, Bad Vöslau (Austria). F. Snyderifusus prevosti (Hörnes, 1853), NHMW 1851/0006/0003, lectotype, Vienna/ Grinzing (Austria). G. Cryptoneolatirus bellardii (Hörnes, 1854), NHMW 1865/0001/0211, Lăpugiu de Sus (Romania). H. Neolatirus recticauda (Fuchs in Karrer, 1877), holotype, NHMW 1883/0002/4168, Rauchstallbrunn-Graben at Baden (Austria). I. Turrilatirus moravicus (Hoernes & Auinger, 1890), NHMW 1866/0001/0860e, paralectotype, Niederleis (Austria). J. Angustifusus hoessii (Naumann,1852), NHMW2023/0350/0006, BadVöslau (Austria).K. Angustifususvindobonensis (Hoernes & Auinger, 1890), NHMW 2023/0338/0020, Lăpugiu de Sus (Romania). L. Fraudifusinus pseudocrispoides nov. sp., NHMW 1868/0001/0462, paratype, Lăpugiu de Sus (Romania). M. Fraudifusinus crispoides (Kittl, 1887), NHMW 2023/0350/0013, Bad Vöslau (Austria). N. Pseudofusus rostratoides nov. sp., NHMW 1864/0001/0204, paratype, Gainfarn (Austria). O. Pseudofusus balteus nov. sp., NHMW 2023/0370/0001, holotype, Letkés (Hungary). P. Pseudofusus stahlschmidti nov. sp., NHMW 1846/0037/0264, holotype, Baden (Austria). Q. Pseudolatirus bilineatus (Naumann, 1852), NHMW 2023/0338/0022, Lăpugiu de Sus (Romania). R. Takashius vinculum nov. sp., NHMW 2023/0348/0008, paratype, Steinebrunn (Austria). S. Takashius boettgeri (Kovács, 2022), NHMW 2023/0338/0023, Lăpugiu de Sus (Romania).

FIGURE 5. Aurantilaria pyrulaeformis (Hoernes & Auinger, 1890). A1–A2. NHMW 1860/0040/0177, lectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1866/0040/0301a, paralectotype, Lăpugiu de Sus (Romania). C. NHMW 1857/0024/0021, paralectotype, Lăpugiu de Sus (Romania). D1–D2. Aurantilaria rudis (Schaffer, 1912), NHMW 1861/0050/0289, lectotype, Gauderndorf (Austria). E1–E2. Aurantilaria rudis (Schaffer, 1912), Collection Anton Breitenberger, Mörtersdorf (Austria).