Aulieezidium karatauense, Szwedo, Jacek & Żyła, Dagmara, 2009

Szwedo, Jacek & Żyła, Dagmara, 2009, New Fulgoridiidae genus from Upper Jurassic Karatau deposits, Kazakhstan (Hemiptera: Fulgoromorpha: Fulgoroidea), Zootaxa 2281, pp. 40-52 : 43-46

Aulieezidium karatauense sp. n.

( Figs. 1–9 )

Diagnosis. Tegmen 3.1 times as long as wide. Cell C5b narrowed in apical portion, closed with a very short icu veinlet, cell C5c with a few veinlets; cell C3 about twice as long as adjoining apical cell; second (apical) veinlet r-m oblique, veinlet im between M1+2 and M3 placed apicad. Other features as for the genus, as it is the only included species.

Description. Total length c. 14 mm. Head of general Fulgoroidea type, lateral margins of frons distinct, postclypeus with lateral carinae being continuation of lateral margins of frons, converging, but not reaching apical portion of postclypeus. Length of head (upper line, lateral view) c. 1.05 mm, length of frons (lateral view) c. 1.87 mm, postclypeus c. 1.35 mm long, anteclypeus c. 1.5 mm long. Rostrum very long (c. 8.3 mm), subapical segment c. 4.3 mm long, apical segment 3.8 mm long, not exceeding length of body.

Pronotum (upper line, lateral view) c. 0.75 mm long, with transverse (postocular) carina present. Mesonotum (upper line, lateral view) c. 1.75 mm long, probably with five longitudinal carinae.

Fore and mid legs normally developed, not flattened. Fore femur 2 mm long, fore tibia 2.75 mm long, fore tarsus c. 0.83 mm long, basitarsomere slightly longer, c. 0.33 mm, mid and apical tarsomeres subequal, c. 0.26 mm long. Mid femur 2 mm long, mid tibia 2.75 mm long, mid tarsus c. 0.83 mm long.

Length of tegmen 11.4 mm, width of tegmen 3.7 mm. Costal margin slightly thickened, stems CA and Pc+CP merely shifted, costal area very narrow, less than half of width of costal cell. Basal cell not distinctly visible but it seems that stems Sc+R and M leave basal cell at same point. Stem Sc+R forked distinctly basad of stem CuA first forking, branch Sc+RA subparallel to costal margin, branch RP forked apicad of stem M branching. Stem M forked at level of claval apex, branch M3+4 forked slightly basad of branch M1+2 forking. Stem CuA forked slightly apicad of stem Sc+R forking, branch CuA1 not forked, branch CuA2 forked basad of claval apex, at level of second veinlet m-cua connection with branch CuA1; branch CuA2a not forked to the tegmen margin; branch CuA2b forked slightly basad of level of M forking, branch CuA2a1 forked again at level of claval apex. Clavus long, exceeding ½ of tegmen length, claval veins Pcu and A1 fused in apical portion of clavus, slightly apicad of branch CuA2 forking. Costal cell with number of elongately sigmoid veinlets. Cell C1 (outer anteapical cell) long, narrower than costal cell, without transverse veinlets basad of claval apex. Anterior discal cell very long without transverse veinlets. Cell C3 twice as long as adjoining apical cell. Posterior discal cell widest, cell C4 long, delimited by two elongately sigmoid veinlets m-cua, the first more oblique, the second partly parallel to longitudinal veins another veinlet m-cua present apicad of claval apex. Cell C5d with a few veinlets, cell C5c narrowed in apical portion, closed with a very short icu veinlet.

Two round spots in apical portion of tegmen, darker marking portion on membrane, two darker markings in median portion of tegmen. Veins light, with darker emarginations.

Etymology. The specific epithet is derived from the name of the Karatau Lagerstätte.

Age and occurrence. Upper Jurassic, Oxfordian–Kimmeridgian, Karabastau Formation. Aulie near Mikhailovka village, Karatau Range, Southern Kazakhstan province, Kazakhstan.

Type material. Holotype, male (?). Specimen number ZPAL V32.354, deposited in the Institute of Palaeobiology, Polish Academy of Sciences, Warsaw, Poland. Collected by Piotr Wegierek.

Figures

FIGURE 1, 2. Aulieezidium karatauense gen. sp. n. 1 — anterior part of body, lateral view; 2 — tegmen venation (partly reconstructed). Scale bar 1 mm.

FIGURE 3 – 8. Aulieezidium karatauense gen. sp. n. 3 — holotype; 4 — tegmen; 5 — apical portion of cells C 5 b and C 5 c; 6 — fragment of costal area and costal cell; 7 — anterior portion of body; 8 — head.

FIGURE 9. Aulieezidium karatauense gen. sp. n. — nomenclature of tegmen veins and cells (modified after interpretations of Emeljanov 1977, Mead & Kramer 1982, Dworakowska 1988, Anufriev & Emeljanov 1988, and Bourgoin 1993).