Augiades bouddha niitakana Sonan, 1936

Chiba, Hideyuki, Hsu, Yu-Feng, Tsukiyama, Hiroshi, Lo, Philip Yik-Fui, Chen, Chien-Ren & Wang, Shou-Ming, 2009, Jinhaku Sonan’s skipper type collection deposited at Taiwan Agricultural Research Institute (Lepidoptera: Hesperiidae), Zootaxa 2202, pp. 48-58 : 53

Augiades bouddha niitakana Sonan, 1936

Zephyrus 6:213.

Labels: “ Augiades / bouddha / niitakana Sonan/ DET. J. SONAN,” “ 800- SUIGEN/ 24 VII 1930 / Col. J. Sonan,” “ No. 60,” “ Type [in red].”

A male specimen was designated as the holotype by Sonan (1936), and this holotype was retrieved in TARI ( Fig. 16–18 ). Evans (1949) treated this taxon as a subspecies of Ochlodes siva Moore, 1878 , and this treatment was followed by Bridges (1983). Shirôzu (1960), however, considered this taxon a synonym of Augiades sylvanus var. formosana Matsumura, 1919 , which was recognized to represent a valid subspecies of Ochlodes subhyalina (Bremer & Grey, 1853) by Evans (1949). Based on the characteristics of male genitalia, Chiba and Tsukiyama (1996) subsequently raised formosana to species status, and considered it a species endemic to Taiwan. Hsu et al. (2006), however, found that formosana Matsumura, 1919 is actually a junior subjective synonym of Ochlodes venata (Bremer & Grey, 1853) , so niitakana Sonan, 1936 should be resurrected in the combination Ochlodes niitakanus . Shirôzu (1960) claimed that the allotype female of niitakana was probably a different species. His statement seems to be groundless. We examined the female paratype of niitakana ( Fig. 19–21 ) and confirmed that it was conspecific with the male holotype.

Figures

FIGURES 1 – 18. Types of skipper taxa described by Jinhaku Sonan: 1 – 3, holotype of Lobocla kodairai, upperside, underside, labels; 4 – 6, holotype of Coladenia sadakoe, upperside, underside, labels; 7 – 9, holotype of Tagiades menaka kotoshona, upperside, underside, labels; 10 – 12, holotype of Notocrypta arisana, upperside, underside, labels; 13 – 15, holotype of Ampittia maro matsumurai, upperside, underside, labels; 16 – 18. holotype of Augiades bouddha niitakana, upperside, underside, labels.

FIGURES 19 – 39. Types of skipper taxa described by Jinhaku Sonan: 19 – 21, paratype of Augiades bouddha niitakana, upperside, underside, labels; 22 – 24, holotype of Parnara kotoshona, upperside, underside, labels; 25 – 27, holotype of Parnara ranrunna, upperside, underside, labels; 28 – 30, holotype of Pamara kiraizana, upperside, underside, labels; 31 – 33, holotype of Gangara thyrsis hainana, upperside, underside, labels; 34 – 36, holotype of Telicota palmarum hainanum, upperside, upperside, labels; 37 – 39, holotype of Tagiades menaka hainana, upperside, underside, labels.