Architipula seebachi ( Geinitz, 1884 )

Kopeć, Katarzyna, Krzemiński, Wiesław, Skowron, Karina & Coram, Robert, 2017, The genera Architipula Handlirsch, 1906 and Grimmenia Krzemiński and Zessin, 1990 (Diptera: Limoniidae) from the Lower Jurassic of England, Palaeontologia Electronica 15 (32), pp. 1-7 : 3

Architipula seebachi ( Geinitz, 1884)

Figures 1 , 2

Synonymy. Phryganidium (Hydropsyche) seebachi Geinitz, 1884 ; Architipula seebachiana Handlirsch, 1906 (new synonymy).

Material examined. SGWG 119 /6, holotype of Architipula seebachi ( Geinitz, 1884) and SGWG 122 /82, holotype of Architipula seebachiana Handlirsch, 1906 . Both from Toarcian ( Lower Jurassic ) of Dobbertin ( Germany) and deposited in the collection Ernst-Moritz-Arndt-Universität Greifswald .

Redescription. Vein Sc ends before mid-point of vein R 2+3+4; Rs long, subequal to veins R 2+3+4 +R 2+3 and R 3 combined; R 4 three times longer than R 2+3+4; stigma large and distinct; d-cell characteristically deeply indented in basal part, upper part of d-cell 1.5- 2 times longer than its lower part (from the forking of Mb to cross-vein m-cu); d-cell about 0.14 the length of the wing; cross-vein m-m between M 1+2 and M 3 distinctly inclined, its bottom part shifted towards the wing base; vein M 3 usually distinctly sinuous; M 1 equal in length to d-cell; cross-vein m-cu situated in the fork of M 3+4 into M 3 and M 4; A 2 usually short and straight.

Remarks. Originally considered to be the type species of the genus, Architipula seebachiana Handlirsch, 1906 is synonymous with Architipula seebachi ( Geinitz, 1884) , which was described as Phryganidium (Hydropsyche) seebachi Geinitz, 1884 and incorrectly assigned to the order Trichoptera , before being transferred to Architipula by Handlirsch in 1906. Both specimens came from the same Toarcian locality of Dobbertin in Germany, and their wings are almost indistinguishable in vein lengths and proportions. Observable morphological differences, such as the degree of curvature of M 4 and the slightly longer Sc vein in A. seebachi , we consider to be accounted for by intraspecific variation.

Figures

FIGURE 1. Wing venation of Architipula seebachi (Geinitz, 1884). SGWG 122/82, holotype of A. seebachiana Handlirsch, 1906.

FIGURE 2. Wing venation of Architipula seebachi (Geinitz, 1884). SGWG 119/6, holotype.