Savignia

Kropf, Holger Frick Wolfgang Nentwig Christian, 2010, Progress in erigonine spider phylogeny-the Savignia-group is not monophyletic (Araneae: Linyphiidae), Organisms Diversity & Evolution (New York, N. Y.) 10 (4), pp. 297-310 : 307

Savignia and Diplocephalus

Clade 44 unites Savignia and Diplocephalus , the two most diverse genera in the Savignia -group. This clade is not well supported, as it relies on a Bremer value of 1 only. Still, out of the two characters that support this node, the AME-lobe (character 106, ci=0.50) is very rarely found. This is a cephalic lobe that only bears the anterior median eyes and can vary from relatively small (in Diplocephalus cristatus ; fig. 39g in Roberts 1993) to very large (in Savignia frontata ; fig. 39f in Roberts 1993). The second occurrence of this character in the current dataset is at the base of the “ Savignia -group” in D. latifrons , which has a very tiny, yet visible AME-lobe (fig. 39i in Roberts 1993; pl. 6H, I in “Electronic Supplementary Material” 2). This lobe is not found very often in erigonine spiders compared to the much more common PME-lobe (e.g. pls. 7H, 9H, 10H, 15H, 18H in “Electronic Supplementary Material” 2; figs. 32–35 in Hormiga 2000). In the current analysis it is only present in two Diplocephalus species and in Savignia . Based on our results it is not possible to judge whether it has a convergent origin or has been reduced and regained in clade 44. The AME-lobe is also found in species that were assigned to other genera (e.g. in Araeoncus galeriformis ; Tanasevitch 1987: fig. 123) but were not included in the present analysis. Its origin should be discussed based on a denser taxon sampling of the Savignia -group, since Diplocephalus was used as a ‘waste’ container for taxa with clear assignment to the Savignia -group but of uncertain placement. Diplocephalus picinus and D. latifrons show all synapomorphies of the Savignia -group but differ from D. cristatus in several characters of the embolic division.