Aplus subpusillus (Hoernes & Auinger, 1890)

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 34-35

Aplus subpusillus (Hoernes & Auinger, 1890)

Figs 15B , 17A–C

* Pollia subpusilla nov. form.—Hoernes & Auinger 1890: 242, pl. 28, figs 12–13.

P [ollia] subpusilla — De Gregorio 1885b: 140.

Pollia (Engina) subpusilla Hö. Au. — Boettger 1906: 32.

Pollia subpusilla R.H. I Auing. — Friedberg 1912: 186, pl. 11, fig. 1.

Cantharus (Pollia) subpusilla Hörnes-Auinger — Csepreghy-Meznerics 1950: 50.

C [antharus]. (P [ollia]). subpusillus (R. Hörn. et Au.) — Sieber 1958: 151.

Cantharus (Pollia) lapugyensis (Hoern. & Auing.) — Báldi 1960: 67, pl. 2, fig. 7 [non Aplus lapugyensis (Hoernes et Auinger, 1890) ].

Cantharus (Pollia) pusillus subpusillus Hoernes & Auinger, 1891 — Strausz 1966a: 307, fig. 139.

Euthria subpusillus R. Hoernes et Auinger, 1879 — Zelinskaya et al. 1968: 192, pl. 45, figs 22–23.

Cantharus (Pollia) aquitanensis Peyr. — Csepreghy-Meznerics 1969: 85, pl. 3, figs 20–21 [non Aplus aquitanensis ( Peyrot, 1928) ].

Cantharus (Pollia) subpusillus Hoernes et Auinger — Csepreghy-Meznerics 1972: 28, pl. 11, fig. 13.

Cantharus (Pollia) aquitanensis Peyr. — Csepreghy-Meznerics 1972: 28, pl. 11, figs 17–18 [non Aplus aquitanensis ( Peyrot, 1928) ].

Cantharus subpusillus (Hoernes & Auinger, 1890) — Bałuk 1995: 241, pl. 37, fig. 4.

Aplus subpusillus (Hoernes et Auinger, 1885) — Kovács & Vicián 2023: 250, figs 12D–E.

non Cantharus (Pollia) subpusillus R. Hoernes et Auinger, 1890 — Iljina 1993: 96, pl. 13, figs 1–2 [= Aplus sinzovi ( Bidzinashvili, 1975) ].

Type material. Lectotype (designated by De Gregorio 1885b: 140), NHMW 1859 /0045/0123, SL: 9.3 mm, MD: 5.7 mm, Niederleis ( Austria), illustrated in Hoernes & Auinger (1890: pl. 28, fig. 13), Figs 17A 1 –A 2 . NHMW 1866 /0001/1066, SL: 10.8 mm, MD: 6.1 mm, Forchtenau ( Austria), illustrated in Hoernes & Auinger (1890: pl. 28, fig. 12), Figs 17B 1 –B 2 , 15B . NHMW 1863 /0015/0465, SL: 11.0 mm, MD: 6.2 mm, Forchtenau ( Austria), Fig. 17C .

Additional paralectotypes. 2 spec., NHMW 1866 /0001/0849, Niederleis ( Austria) . 35 spec., NHMW 1863 /0015/0668, Niederleis ( Austria) . 26 spec., NHMW 1869 /0001/0484, Forchtenau ( Austria) .

Revised description. Small, squat biconic shell of up to five teleoconch whorls; apical angle 55–62°. Protoconch low conical of 2.5 convex whorls, diameter: 700 μm, height: 700 μm. Early teleoconch whorls almost flat-sided, periphery at abapical suture, with narrow, deeply incised suture. Sculpture of three close-set spiral cords strongly swollen almost tubercular over ribs. Single secondary spiral cord intercalated between adapical and mid-whorl primary spirals on third teleoconch whorl. Additional secondary spiral appears on penultimate whorl at abapical suture. Last whorl attaining 65–67% of total height, strongly convex with rounded periphery and conical base, bearing 11 axial ribs and ten elevated, rounded spiral cords; one secondary spiral cord intercalated in each interspace, secondary between two adapical primaries more prominent than others.Aperture moderately wide, ovate. Columella excavated in upper half. Columellar callus forming broad, thickened rim, sharply delimited from base, with about four weak to subobsolete columellar folds, angled at transition to siphonal canal, accentuated by abapical columellar fold. Anal canal broad U-shaped, moderately incised in thickened adapical tip of aperture, accentuated by small parietal denticle and larger anal denticle. Outer lip strongly thickened by terminal varix, with seven elongated denticles almost extending to peristome. Siphonal canal moderately short, wide, slightly deflected to the left, weakly notched.

Discussion. This species is characterized by its small size, squat biconic outline and tubercular sculpture. The superficially similar but larger Aplus moravicus (Hoernes & Auinger, 1890) differs in its convex spire whorls and prominent lirae inside the outer lip. Moreover, it has a longer siphonal canal. This species was confused by Csepreghy-Meznerics (1969, 1972) with Aplus aquitanensis ( Peyrot, 1928) from the Early Miocene of France. Aplus aquitanensis differs from Aplus subpusillus (Hoernes & Auinger, 1890) in its higher last whorl and narrower aperture (see Peyrot 1927: pl. 4, figs 40–41; Lozouet 2021: pl. 40, fig. 9).

Iljina (1993) listed Aplus subpusilla from the Konkian of the Mangyshlak Peninsula ( Kazakhstan). The illustrated Eastern Paratethyan specimens, however, differ considerably from the Central Paratethyan species in their prominent axial ribs, weak spiral sculpture and weaker denticles in the inner lip. This species was described by Bidzinashvili (1975) as Genota sinzovi and therefore, we propose Aplus sinzovi ( Bidzinashvili, 1975) as new combination for it.

Paleoenvironment. Probably inner neritic. At the Niederleis section, sediment and fossils from coastal and lagoonal environments not exceeding 30 m water depth were transported by tempestites into offshore settings ( Mandic et al. 2002).

Distribution in Central Paratethys. Badenian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin: Zboriv (= Zborów) ( Ukraine) ( Friedberg 1912); Korytnica Basin: Korytnica ( Poland) ( Bałuk 1995); Vienna Basin: Niederleis ( Austria); Boršov (= Porstendorf), Porzteich at Břeclav ( Czech Republic) (Hoernes & Auinger 1890); Eisenstadt-Sopron Basin: Forchtenau ( Austria) (Hoernes & Auinger 1890); North Alpine-Carpathian Foreland Basin: Lysice ( Czech Republic) (Hoernes & Auinger 1890); Pannonian Basin: Hidas, Letkés, Szokolya ( Hungary) ( Báldi 1960; Strausz 1966a; Kovács & Vicián 2023); Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972); Făget Basin: CoŞteiu de Sus ( Romania) ( Boettger 1906).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 15. Protoconchs of Paratethyan Pisaniidae, Prodotiidae and Tudiclidae. A. Aplus moravicus (Hoernes & Auinger, 1890), NHMW 1862/0029/0034, lectotype, Rudice u Blanska (Czech Republic). B. Aplus subpusillus (Hoernes & Auinger, 1890), NHMW 1866/0001/1066, paralectotype, Forchtenau (Austria). C. Aplus transsylvanicus (Hoernes & Auinger, 1890), NHMW 1873/0026/0019b, lectotype. D. Aplus zebus (De Gregorio, 1885a), NHMW 1855/0045/0439, Gainfarn (Austria). E. Aplus anatolicus nov. sp. RGM 784 005, paratype, Seyithasan (Turkey). F. Hilda transsylvanica (Hoernes & Auinger, 1884), NHMW 1876/0011/0023a, lectotype, Lăpugiu de Sus (Romania).G. Janiopsis vindobonensis nov. sp., NHMW 1850/0009/0002, holotype, Baden (Austria). H. Monostiolum hungaricum (Csepreghy-Meznerics, 1954), NHMW 1868/0001/0094, Steinebrunn (Austria). I. Dianthiphos klaudiae (Kovács & Vicián, 2023), NHMW1846/0037/0144a, Steinebrunn (Austria). J. Euthria obelixi nov. sp., NHMW 1865/0015/0060, holotype, Lysice (Czech Republic). K. Tethyspollia mariae (Hoernes & Auinger, 1890), NHMW 1868/0001/0441, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 17. Aplus subpusillus (Hoernes & Auinger, 1890). A1–A2. NHMW 1859/0045/0123, lectotype, Niederleis (Austria). B1–B2. NHMW 1866/0001/1066, paralectotype, Forchtenau (Austria). C. NHMW 1863/0015/0465, paralectotype, Forchtenau (Austria).