Anthobium puetzi, Shavrin & Smetana, 2017

Shavrin, Alexey V. & Smetana, Aleš, 2017, A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups, Zootaxa 4365 (1), pp. 1-39 : 19-22

3.2.2.3. Anthobium puetzi sp.n.

( Figs. 41 , 58, 61–62 )

Type material examined: Holotype ♂: ‘ China: Sichuan, Ganzi pref. | Daxue Shan , 102.00 30.03 N, | 5 km E Kangding, river valley | ca. 3000m, 20./ 23.V.1997, leg. A. Pütz’ <rectangular label, printed>, ‘HOLOTYPE | Anthobium | puetzi sp.n. | Shavrin A. & Smetana A. 2017’ <red rectangular label, printed> (temporarily in CAP, to be eventually deposited in SNSD).

Paratypes (3 specimens): 1 ♂, 1 ♀: same label as the holotype (CAP); 1 ♂ [plastic plate with aedeagus, abdominal tergite VIII, sternite VIII and apical segment in Canadian balsam was pinned under the plate with beetle]: ‘CHINA: W-Sichuan | Aba Tibetian Aut. Pref. | Weizhou Co., Qionglai Shan | Wolong Valley, 69km WSW | Guanxian, 3900 m, 30.53,67N | 102.54,87E, 15.VII.1999 | leg. A. Pütz’ <rectangular label printed> (CAS). All paratypes with additional red rectangular printed label: ‘PARATYPE | Anthobium | puetzi sp.n. | Shavrin A. & Smetana A. 2017’.

Description. Measurements (n=4): HW: 0.85–0.92, HL: 0.52–0.59; AL (holotype): 2.03; OL: 0.24–0.25; PL: 0.79–0.82; PW: 1.26–1.37; ESL: 1.90–2.00; EW: 1.75–1.97; AW: 1.35–1.55; MTbL (holotype): 1.12, MTrL (holotype): 0.52 (MTrL 1–4: 0.32; MTrL 5: 0.20); AedL: 0.90–0.97; TL: 3.75 (holotype)–4.75.

In external appearance similar to A. morchella sp.n. Body moderately elongated. Reddish-brown to brown; antennomeres 3–11 brown; lateral and basal portions of pronotum, latero-apical and apical portions of elytra, apical segments of abdomen yellow-brown; mouthparts, antennomeres 1 and 2 and legs yellow. Head behind postocular ridges and lateral portions of neck with fine small cellular sculpture, larger and smoothed in middle of neck. Punctation of infraorbital ridges moderately regular, markedly denser than that on middle part of head; middle part of neck with transverse, irregular row of deep large punctures; pronotum with coarse and deep punctures, denser on medio-basal portion, irregular and somewhat smaller on medio-apical part, more rugose on lateral portions; scutellum with several small punctures; punctation of elytra as that on pronotum, slightly denser and smaller in prescutellar and basal area near suture. Habitus as in Fig. 41 .

Head 1.5–1.6 times as wide as long, slightly widened posteriad; middle portion of head and infraorbital ridges slightly elevated; median portion with a pair of indistinct medial impressions behind level of anterior margin of eyes and small transverse impression on level of posterior margin of eyes, with very deep elongated grooves in front of ocelli reaching apical third of length of eye; head gradually narrowed toward neck behind distinctly smoothed postocular ridges; anterior portion between antennal insertion and anterior margin of eye with wide, moderately deep notch. Ocelli situated slightly behind of level of postocular ridge, distance between ocelli slightly larger than distance between ocellus and posterior margin of eye. Apical segment of maxillary palp less than three times as long as preceeding segment. Measurements of antennomeres (holotype): 1: 0.21 × 0.09; 2: 0.15 × 0.06; 3: 0.18 × 0.06; 4–7: 0.19 × 0.06; 8: 0.17 × 0.06; 9–10: 0.17 × 0.07; 11: 0.22 × 0.07.

Pronotum 1.5–1.6 times as wide as long, 1.4 times as wide as head, widest near middle; lateral edge with moderately uniform crenulation, distinctly larger in middle; middle elevation of pronotum with vague, moderately wide longitudinal and a pair of suboval medio-basal impressions; small area above medio-basal impressions markedly elevated; lateral portions narrow, indistinctly explanate.

Elytra more than twice as long as pronotum, gradually widened apicad, reaching basal part of abdominal tergite V–VII; shoulders indistinctly protruded anteriad; middle parts of each elytron with vague irregular elevations between punctures (one paratype with indistinct longitudinal elevation beginning behind shoulders and stretching to apical third of elytra).

Male. Apical margins of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus ( Fig. 58 ) with wide basal part slightly narrowed towards subtruncate apex of median lobe; parameres significantly exceeding apex of median lobe, with moderately long apical and two preapical setae; internal sac narrow and long, markedly more sclerotized basally. Aedeagus laterally as in Fig. 61 .

Female. Apical margins of abdominal tergite VIII and sternite VIII straight to slightly rounded.

Comparative notes. Based on the presence of indistinct irregular elevations between punctures on the elytra and parameres significantly exceeding apex of median lobe of the aedeagus, A. puetzi sp.n. is similar to A. morchella sp.n., from which it differs by the head behind eyes slightly widened posteriad, by the larger and more rugose punctation of the pronotum with vague longitudinal and medio-basal impressions on middle elevation and by the relatively shorter parameres. Anthobium puetzi sp.n. differs from remaining two species of the morchella group by wider median lobe of the aedeagus and by the presence of two long preapical setae on apical portions of parameres.

Distribution. The new species is known from two locations ( Fig. 62 ) in Daxue Shan and Qionglai Shan ranges in Sichuan, China.

Bionomics. Specimens were taken by sifting forest floor litter at elevations from 3000 to 3900 m a.s.l. Etymology. Patronymic, the species is named to honour our colleague Andreas Pütz (Eisenhüttenstadt, Germany), the collector of the type specimens.

Figures

FIGURES 39–41. Habitus of Anthobium morchella group: 39—A. morchella (paratype), 40—A. hydraenoides (paratype), 41—A.puetzi (holotype).Scale bars:1.0 mm.

FIGURES 56–61. Aedeagus of Anthobium morchella (56, 59—paratype), A. hydraenoides (57, 60—paratype) and A. puetzi (58, 61—paratype): 56–58—parameral view, 59–61—lateral view.Scale bars: 0.1 mm.

FIGURE62.Distribution of Anthobium morchella group inChina:A.morchella(black square), A.hydraenoides(black circle), A. puetzi (black triangle).