Anomala

Filippini, Valentina, Micó, Estefania & Galante, Eduardo, 2013, Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae), Zootaxa 3670 (2), pp. 255-273 : 270-271

Key to dorsally setose Anomala species of the Neotropical Region

Only the species with setae on the elytral surface and two protibial teeth are included. Of the additional species included in the key, only A. amphicoma probably belongs to the same group of the species described here. A. semitonsa , A. balzapambae , and especially A. hispidula and A. tindakua are likely distant phylogenetically.

1. Rounded shape, with widest point mid length of elytra; two transversal bands on elytra: a median transversal wavy band, hind band usually not reaching the posterior margin of elytra....................................................... 2

-. Elongate shape, with widest point at last third of elytra; elytra with uniform colour, or with 1 or more, continuous, transversal bands, the hind band covering the apical third of elytra; when bands are defined by spots, they are arranged in 3 rows...... 3

2. Median band on elytra continuous, hind band reaching sides, male genitalia illustrated in Fig. 37 . Panama, Ecuador........................................................................................ A. semitonsa Bates, 1888

-. Median band on elytra composed of isolated spots, hind band usually not reaching sides, male genitalia illustrated in Fig. 38 . Costa Rica, Ecuador............................................................ A. balzapambae Ohaus, 1897

3. Pronotum metallic green, larger at 1/3 of height than at base, elytra uniform dark brown, total length about 10 mm, male genitalia illustrated in Fig. 39 . Panama................................................... A. amphicoma Bates, 1888

-. Pronotum dark brown, largest at base, if a green luster is present then elytra have transversal light bands................ 4

4. Pronotum almost rectangular, head large, almost as long as pronotum and 2/3 the width of the pronotum. Elytra light brown dappled with darker spots, concentrated in three transversal bands (post humeral, median, and posterior half of elytra), or expanding until covering almost all surface................................................................ 5

-. Pronotum trapezoidal, head small, about half wide as the pronotum. Elytra with uniform colour or maximum two transversal bands, never a posthumeral band........................................................................ 6

5. Elytra with setae on the entire surface, pronotum of uniform colour or with slightly lighter border in light coloured specimens, pattern on elytra not defined, aedeagus illustrated in Fig. 40 . Mexico, Guatemala................ A. hispidula Bates, 1888

-. Elytra with setae only on posterior third, border of pronotum always of lighter colour, pattern on elytra well defined, aedeagus in Morón & Nogueira (2002). Mexico...................................... .. A. tindakua Morón & Nogueira, 2002

6. Elytra uniformly light brown or with one transversal dark band. Size approximately 11 mm. Costa Rica...................................................................................... A. flavacoma Filippini, Micó , & Galante

-. Elytra uniformly dark brown or with two bands, apical third of elytra always dark.................................. 7

7. Pronotum with irregular surface due to small depressions on sides of median sulcus. Parameres wide, maximum length 3/4 of tectum length, with blunt and wide apex. Length less than 10.1 mm. Costa Rica..................................................................................................... A. megaparamera Filippini, Micó , & Galante

-. Pronotum with uniform surface, sometimes a median sulcus is present. Parameres slender, with defined narrow apex; length of parameres not reaching 3/4 of tectum length. Length 8.4–12.7 mm .............................................. 8

8. Elytra uniformly dark or with a lighter posthumeral band (rarely a second light band on disc); male protibia upper tooth short (less than 1/4 of total length) and almost straight; aedeagus with wide apex and strongly sinuate ventral margin. Costa Rica................................................................... A. pseudoeucoma Filippini, Micó , & Galante

-. Two darker transversal bands are visible on elytra; male protibia upper tooth long (more than 1/4 of total length) and oblique, parameres with short rounded apex and slightly sinuate ventral margin. Guatemala, Costa Rica...... A. eucoma Bates, 1888

Figures

FIGURE 37 – 40. 37. a. Aedeagus and b. endophallus of Anomala semitonsa (syntype, Bugaba, Panama, MNHM); c. view from the other side of dorsal sacculus of endophallus, with a flat sclerotized plate in evidence. 38. a. Aedeagus and b. endophallus of Anomala balzapambae (holotype, Balzapamba, Ecuador, MNHUB); c. view from the other side of dorsal sacculus of endophallus, with a flat sclerotized plate evident. 39. a. Aedeagus and b. endophallus of Anomala amphicoma (Volcán de Chiriqui, Panama, MNHN). 40. Aedeagus of Anomala hispidula (syntype, San Geronimo, Guatemala, MNHM). Scale = 1 mm.