Anisochrodes janavlada, Švihla, 2012

Švihla, Vladimír, 2012, A review of the family Oedemeridae (Coleoptera) of the Socotra Archipelago, Acta Entomologica Musei Nationalis Pragae 52, pp. 337-346 : 342-344

Anisochrodes janavlada sp. nov.

( Figs. 6–13 )

Type locality. Yemen, Socotra Island, Wadi Zirik, 12°29.584′N 53°59.475′E.

Type material. HOLOTYPE: J ( NMPC), ‘ YEMEN, Socotra Isl. / Wadi Zirik, 12.vi.2010 / N 12°29,584′, E 053°59,475′ / V. Hula & J. Niedobová leg. [white label, printed]’. PARATYPES ( NMPC), same label data, 1 ♀; ‘ YEMEN, Socotra Isl. / Firmihin plato, 400-500 m / N 12°28′46″, E 54°00′89″ / 18.-19.vi.2010 / V. Hula & J. Niedobová leg. [white label, printed]’, 1 ♀; ‘ YEMEN, Socotra Isl. / Dgisfu valley, 2.vi.2010 / N 12°28,444′ E 054°08,596′ / V. Hula & J. Niedobová leg. [white label, printed]’, 1 ♀; ‘ YEMEN, Socotra Isl. / Zemhon area 270-300 m / N 12°20,58′, E 054°06,39′ / 16.-17.6., V. Hula leg. [white label, printed]’, 1♀; ‘ YEMEN, Socotra I., 4.-5.vi. / Qualentiah env., 2010 / slopes 5 km SE from Quasoit / N12°39,691′, E 053°26,658′ / V. Hula & J. Niedobová leg. [white label, printed]’, 1 ♀; ‘Republic of Yemen / Socotra Isl. – Deigyup cave env. / V. Hula lgt., 16.6.2009 [white label, printed]’ 7 JJ 1 ♀; ‘ YEMEN, Socotra Isl. / Deiqub cave env. / 10.vi.2010 / V. Hula & J. Niedobová leg. [white label, printed]’, 1 J 2 ♀♀; ‘ YEMEN, Socotra Isl., 2.vi.2010 / 7 km NW from Rhi di-Hamri, / flowering Croton scrubs / V. Hula & J. Niedobová leg. [white label, printed]’, 12 JJ 3 ♀♀; ‘ SOCOTRA Is. ( YE) / Noged plain (sand dunes) / SHARET HALMA vill. env. / 12°21,9′N, 54°05,3′E, 20 m / Jan Batelka leg. 10-11.xi.2010 [white label, printed]’, 1J; ‘ YEMEN, SOCOTRA ISLAND / WADI THAR DI ITRUR, 21.vi.2012 / sifting under Ficus cordata / 12°32.8′N 53°42.9′E, 54 m // SOCO- TRA expedition 2012 / J. Bezděk, J. Hájek, V. Hula / P. Kment, I. Malenovský / J. Niedobová & L. Purchart leg.’, 1 J; ‘ YEMEN, SOCOTRA ISLAND / ALOOVE AREA, Aloove vill. env. / Jatropha unicostata shrubland with / Boswellia elongata trees, / 19.–20.vi.2012 / 12°34.5′N 54°07.4′E, 221 m // SOCOTRA expedition 2012 / J. Bezděk, J. Hájek, V. Hula / P. Kment, I. Malenovský / J. Niedobová & L. Purchart leg.’, 6 JJ 3 ♀♀; ‘ YEMEN, SOCOTRA ISLAND / HOMHIL protected area / open woodland with Boswellia & Dracaena trees, 10.–11.vi.2012 / 12°34.5′N 54°00.9′E, 115 m // SOCOTRA expedition 2012 / J. Bezděk, J. Hájek, V. Hula / P. Kment, I. Malenovský / J. Niedobová & L. Purchart leg.’, 4 JJ 2 ♀♀; ‘ YEMEN, SOCOTRA ISLAND / DEIQUB cave, 12.vi.2012 / cave & Croton socotranus + / Jatropha unicostata shrubland / 12°23.1′N 54°00.9′E, 115 m // J. Bezděk, J. Hájek, V. Hula / P.Kment, I. Malenovský / J. Niedobová & L. Purchart leg.’, 1 J 2 ♀♀; ‘ YEMEN, SOCOTRA ISLAND / KAZAZHAN area / shrubland on limestone; sifting / 10.vi.2012 / 12°33.8′N 54°19.8′E, 540 m // SOCOTRA Expedition 2012 / J. Bezděk, J. Hájek, V. Hula, / P. Kment, I. Malenovský, / J. Niedobová & L. Purchart leg.’, 2 JJ 1 ♀.

Description. Coloration ( Figs. 6–8 ). Head iron grey, sometimes with slight coppery tinge, mouthparts terra-cotta, tips of mandibles sepia, antennae terra-cotta. Pronotum either entirely terra-cotta or with pair of smaller or larger spots in anterior half or spots enlarged and extending over almost whole length of pronotum, so that only narrow margins of pronotum remain yellow. Legs terra-cotta, tibiae mostly longitudinally infuscate on outer side, tarsi sepia. Ventral part of abdomen iron grey, sometimes with slight coppery tinge, last two abdominal segments terra-cotta. Elytra iron grey.

Male. Eyes rather small, slightly reniform, moderately protruding, head across eyes only very slightly wider than pronotum, head behind eyes narrowing posteriorly almost in straight line. Antennae filiform, moderately exceeding basal third of elytral length, terminal antennomere constricted behind its midlength. Terminal palpomere of maxillary palpus securiform. Surface of pronotum finely and densely imbricate-punctate, with fine, white, recumbent pubescence, matt. Pronotum about as long as wide, moderately cordiform, anterior margin very slightly rounded, anterior corners rounded, lateral margins sinuately converging posteriorly, posterior corners rounded, posterior margin straight. Surface of pronotum more finely and sparsely imbricate-punctate than that of head and with yellow pubescence, semilustrous. Basal projections of claws long, almost reaching apices of claws. Elytra parallel-sided up to elytral midlength, moderately narrowing in posterior half, elytral apices separately rounded. Elytral venation only very slightly visible basally, surface of elytra very finely and densely rugulose-lacunose, with fine, white, short recumbent pubescence, matt. Pygidium almost twice as long as apical sternite, apex of which is widely rounded with shallow, subtriangular emargination in middle; pygidium subtriangular with shallow, subtriangular emargination apically ( Fig. 10 ). Projections of urite VIII visible, very narrow, moderately curved, similar to that of A. jelineki Švihla, 1983 (cf. Fig. 10 ). Tegmen and aedeagus as in Figs. 11–13 .

Female. Basal projections of claws shorter than in male, hardly reaching midlength of claws, elytral venation better developed, very slight but visible in ca basal two thirds of elytral length. Pygidium only moderately exceeding last sternite, similar to that of male, last sternite subtriangular, roundly tapering apically from about its midlength ( Fig. 9 ).

Length (J ♀): 4.8–11.9 mm.

Differential diagnosis. Anisochrodes janavlada sp. nov. with its pronotum at least partly coloured differently than elytra mostly resembles A. escalerai Švihla, 1999 , from which it differs in much finer and sparser pubescence of elytra, semilustrous pronotum, unicolor tibiae and especially in parameres with basoventral tooth and bigger apical hook of the aedeagus (cf. Figs. 6–8 and ŠVIHLA 1999). The three remaining species of the genus have pronotum similar to elytra in colour, and denser and thicker elytral pubescence. This type of pubescence suggests that the new species represents a sister clade of the three species known from the Asian mainland.

Etymology. Patronymic, dedicated to two of its collectors, Jana Niedobová (Brno, Czech Republic) and Vladimír (familiarly Vláďa) Hula (Moutnice, Czech Republic). To be used as the noun in apposition.

Distribution. Endemic to Socotra Island.

Figures

Figs.5–8. General habitus of Socotran Oedemeridae. 5 – Dentostomus guichardi (Švihla, 1987); 6–8 – Anisochrodes janavlada sp. nov.

Figs.9–13. Anisochrodes janavlada sp. nov. 9 – last abdominal segment of female, ventral view; 10 – last abdominal segment of male including projections of urite VIII, ventral view; 11 – tegmen, ventral view; 12 – apical portion of tegmen, lateral view; 13 – aedeagus, lateral view. Scale bar: 0.5 mm.