Anarrhinum fruticosum subsp. demnatense (Cosson) Maire, 1934

Ferrer-Gallego, P. Pablo, 2021, Typification of the accepted names in the variable species Anarrhinum fruticosum Desf. (Antirrhineae, Plantaginaceae), Adansonia (3) 43 (4), pp. 31-36 : 34

Anarrhinum fruticosum subsp. demnatense (Cosson) Maire

Catalogue des plantes du Maroc, vol. 3: 681 (1934).

LECTOTYPIFICATION

Sutton (1988: 257) indicated the type as: “Type: Morocco: Ga; Demnate, Djebel Imiifry, 18 vii 1879, Ibrahim s.n. (iso. BM!)”. However, this is not an effective designation according to Art. 9.12 of the ICN because this specimen at BM cannot be considered as part of the exsiccatum of the Société Dauphinoise cited in the protologue.In fact, Cosson (1891: 84), in the protologue, stated: “ A. Demnatense Coss. in herb. – Exs. Soc. Dauph. Mar. 522 bis (sub A. fruticosum )”. A diagnosis, a complete description of the species is provided by Cosson (l.c.), as well as the provenance “In collibus calcareis apricis. – Mar. Dj. Imiifry in provincia Demnate ad boreali-orientem urbis Maroc. Imperio Maroccano peculiare”. Moreover an excellent and detailed illustration was given (Tab. 152, showing two stems with leaves, flowers and fruits, and several flowers details; image available at https://www.biodiversitylibrary.org/item/113581#page/53). The specimens cited by Cosson (1891) in the protologue are syntypes according to the Art. 9.6 and Art. 40 Note 1 of ICN. Based on the Art. 9.12 of the ICN, this material has priority in the lectotype designation.

There is a specimen at P ( P038882611 ) which bears six stems, with leaves, flowers and fruits, and a printed label (“ Société dauphinoise, 1882. / Nº 522 bis. Anarrhinum fruticosum / Desf. fl. Atl. II, p. 52, t. 142. – DC. Prodr. X, 289. / Djebel Imiifry, province de Demnat ( Maroc). / 4 juillet 1881. / Leg. Ibrahim, / Dr. E. Cosson.”) ( Fig. 2 ). In addition, another specimen at P ( P03882610 ) was traced. Although not identified as “Société dauphinoise, 1882/Nº 522 bis ”, it belongs probably to this collection since it was collected in the same place by Ibrahim on 4 July 1881, and likely to be a syntype. On the other hand, other specimens (see e.g., P [barcode P03882612 ] and MPU [barcode MPU237886 ; image available at https://explore.recolnat.org/occurrence/ 2314BEC16FA04046B07B81C98F8A9F8C]) were collected by Ibrahim in Djebel Imiifry (Demnate), but on 18 July 1879 (as the specimen at BM cited by Sutton [1988] as “type”), and not on 4 July 1881 as reported in the above mentioned syntypes ( P038882611 and P03882610 ) .

In conclusion, among the material above cited, I here designated as the lectotype of Anarrhinum demnatense the specimen P03888261 because it is the only sure specimen that can be identified as part of the exsiccatumof the Société Dauphinoise with number “ 522 bis ”, such as part of the original material (Art. 9.4 of ICN).

Figures

FIG. 2. — Lectotype of Anarrhinum demnatense Cosson (P038882611).