Amplirhagada inusitata, Köhler, 2011

Köhler, Frank, 2011, Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae), Records of the Australian Museum 63 (2), pp. 167-202 : 185-186

publication ID

https://doi.org/ 10.3853/j.0067-1975.63.2011.1581

publication LSID

lsid:zoobank.org:pub:1BCD4085-D2B9-400D-B504-8C85C30303D6

persistent identifier

https://treatment.plazi.org/id/5E55C8D9-5E35-4584-8FF0-EC0DE7DB76B4

taxon LSID

lsid:zoobank.org:act:5E55C8D9-5E35-4584-8FF0-EC0DE7DB76B4

treatment provided by

Felipe

scientific name

Amplirhagada inusitata
status

sp. nov.

Amplirhagada inusitata View in CoL n.sp.

Type locality. Western Australia, NW Kimberley, N side of Prince Frederick Harbour , near mouth, 16 km W of Mt. Anderdon, 0.3 km from coast; 14°57'10"S 125°16'30"E ( RFS 14-1 ; coll. V. Kessner, 14 Jun 1987) ( Fig. 1 View Figure 1 ) GoogleMaps .

Type material. Holotype WAM S34728 View Materials (preserved specimen, dissected) . Paratypes WAM S34729 View Materials (preserved specimen, juvenile) , FMNH 220537 About FMNH (preserved specimen) , AM C.472929 (2 dried shells), WAM S34730 View Materials (3 dried shells) (Pl. 1.9) , FMNH 220540 About FMNH (6 dried shells) .

Etymology. Species epithet derived from “inusitata” (Latin, unusual, uncommon, extraordinary; of female gender) in reference to its remarkably distinct penial morphology.

Description

Shell (Pl. 1.9; Fig. 26 A–C View Figure 26 ). Broadly conical with low spire. Thin, translucent. Periphery sharply angulate; upper sector of whorls flattened. Umbilicus 30–50% concealed by columellar reflection. Background colour pale yellowish brown; peripheral band diffuse to well developed, moderately broad, brown, visible on visible on last whorl only; subsutural band diffuse to well developed, brown, visible on last whorl only; outer and inner colour like shell. Protoconch 2.6 mm in diameter, comprising about one whorl, with very indistinct radially elongated pustulations. Teleoconch with faint axial growth lines, evenly distributed across shell surface. Angle of aperture 45–60 degrees; outer lip thin, well rounded, slightly expanded; basal node absent or weak. Parietal wall of inner lip inconspicuous. Average shell size 16.8±1.7 × 21.4± 0.7 mm ( Table 1).

Radular and jaw morphology ( Fig. 26 D–G View Figure 26 ). Tooth formula C + 14–19 + 4 +?; 125 rows of teeth (n = 1). Jaw with 10 plates.

Genital morphology ( Figs 27–28 View Figure 27 View Figure 28 ). Penis long, curved to bending, tubular, clearly longer than anterior part of oviduct. Penial sheath very delicate. Penial retractor muscle very short. Penial verge extremely elongated, extending almost entire length of penis, with pointed tip, proximally the lumen of the penial verge forms an extended duct that is connected to vas deferens, possibly functioning as epiphallus. Penial chamber extending only distal half of entire length of penial complex, inner penial wall almost entirely smooth, in proximal third corrugated longitudinal pilasters are present consisting of fused pustules. No main stimulatory pilaster differentiated. Vas deferens rather thin, undulating, entering penial sheath close to penial apex. Vagina moderately long, tubular to slightly inflated; inner vaginal wall and wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix reaching base of spermoviduct. Free oviduct long, bending, tubular, comprising more than half of length of anterior part of oviduct. Spermoviduct as long as anterior part of oviduct.

Aestivation strategy. Free sealer.

Remarks. Anatomical description based on dissection of the holotype. Listed by Solem (1991) as “ Amplirhagada NSP 26”. Shell similar to other relatively large and broadly conical species with spiral bands, such as A. combeana Iredale, 1938 and A. mitchelliana Solem, 1981 , differing from all other congeners by extremely long and tubular penial verge, narrowly elongated penis, smooth penial wall with only short corrugated pilasters at its base. Found in sympatry with A. pusilla Solem, 1981 and A. epiphallica n.sp. described below.

V

Royal British Columbia Museum - Herbarium

WAM

Western Australian Museum

AM

Australian Museum

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