Amphidromus (Amphidromus) givenchyi Geret, 1912

Inkhavilay, Khamla, Sutcharit, Chirasak & Panha, Somsak, 2017, Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species, European Journal of Taxonomy 330, pp. 1-40 : 14-15

Amphidromus (Amphidromus) givenchyi Geret, 1912

Figs 1 , 2C , 3C , 4I ; Table 1

Amphidromus givenchyi Geret, 1912: 55–56 , pl. 2, figs 21–22. Type locality: Unknown. Amphidromus givenchyi – Laidlaw & Solem 1961: 526, 621. — Richardson 1985: 43. — Sutcharit & Panha 2006: 26–28, figs 4n–q, 18–19.

Material examined

LAOS: 120D + 18D juveniles ( Figs 3C , 4I , CUMZ 7015), 7D specimens, in ethanol ( Fig. 2 C , CUMZ 7016), Thad Lor Waterfall, Salavan District, Salavan; 61D + 8D juveniles ( CUMZ 7017), 2D specimens, in ethanol ( CUMZ 7018), Wat Phou (Temple on the mountain), Khong Xédon District, Salavan.

Remarks

The species description, as well as the morphology of the genitalia and the radula, have been carefully presented in Sutcharit & Panha (2006), which allows an unambiguous recognition of this species. The unique characters are a relatively medium sized (compared to other members of this genus, as shown in Table 1 ), ovate-conical and dextral shell; apex with black spot; aperture oblique; peristome expanded and not reflected; parietal callus thin and transparent to slightly thickened and white; shell color uniform greenish with pale to yellowish color on earlier whorl, and greenish subsutural band; genitalia contain long and slender penis, short and curved epiphallus of equal length to flagellum; proximal to enlarged and coiled appendix epiphallus; appendix short.

The early records of this species were from the Ubon Ratchathani and Kalasin Provinces of Thailand, and the Luang Phrabang and Savannakhet Provinces of Laos ( Sutcharit & Panha 2006). Here, two more localities from Salavan Province, Laos are added. All of the records are from dry dipterocarp to deciduous forests with sandstone hills. The snails were found in small holes on branches or tree trunks during the aestivation period ( Fig. 3C ).

Figures

Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study.

Fig. 2. Living snails. A–B. Amphidromus roseolabiatus Fulton, 1896. A. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7012). B. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7005). C. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7018). D. Amphidromus syndromoideus sp. nov., holotype from the type locality (CUMZ 7019).

Fig. 3. Aestivation sites of Amphidromus species. A–B. Amphidromus roseolabiatus Fulton, 1896 (shell height about 34 mm) from NamTurn Bridge, Bolikhamxay, Laos, aestivated. A. Inside sterile fronds of stag horn ferns, Platycerium Desv. B. In a small hole on a Jackfruit tree trunk, Artocarpus heterophyllus Lam., with other snails, Quantula Baker, 1941 and Durgella Blanford, 1863. C. Amphidromus givenchyi Geret, 1912 (shell height about 43 mm), hiding in a small hole of a dipterocarpus tree shell. D. Amphidromus syndromoideus sp. nov. (shell height about 28 mm) from the type locality, hiding in a hole of a tree, visible after removal of clusters of sterile fronds of the basket fern Drynaria (Bory) J. Sm.

Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann & Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020).