Sangabasis feliculoi, Villanueva, R. J. T. & Dow, R. A., 2014

Villanueva, R. J. T. & Dow, R. A., 2014, Review of the Philippine taxa formerly assigned to the genus Amphicnemis Selys, II. Genus Sangabasis with descriptions of eight new species (Odonata: Coenagrionidae), Zootaxa 3815 (1), pp. 1-28 : 15-19

Sangabasis feliculoi sp. nov.

( Figs. 13 , 25 , 35 , 48 , 60 , 65 )

Amphicnemis sp. ( cf. dentifer );— Hämäläinen & Müller (1997: 259, 278: Note 31; part, population from Samar).

Type material (all in RMNH, all Philippines, Samar Island, Western Samar Province). Holotype: ♂, Hinabangan, Bagakay, Arizona, 180–200m, 29 viii–20 ix 1996, leg. A. Buenafe. Paratypes: 20 ♂ Hinabangan, Bagakay, Arizona, 100–200m, 29 viii–20 ix 1996, Leg. A. Buenafe.

Other material (all Philippines, Samar Island, all in RMNH unless otherwise stated). Western Samar Province: ♂, Hinabangan, 50 – 150m, 1–14 iv 1992, leg. T. Borromeo jr; ♂, Hinabangan, 7 iv 1992, leg. T. Borromeo Sr”; 38 ♂, 8 ♀, Hinabangan, Bagakay, Arizona, 100–200m, 29 viii–20 ix 1996, leg. A. Buenafe; 162 ♂, 27 ♀, Hinabangan, Arizona, 100–280m, 29 iii–6 iv 1997, leg. R.A. Müller; 21 ♂, 6 ♀, Hinabangan, Arizona area, 100–280m, 5–12 v 1997, leg. A. Buenafe; 2 ♂, 2 ♀, Hinabangan, San Isidro, San Isidro River 90–200m, 31 iii–5 iv 1997, leg. R.A. Müller; 2 ♂, Basey, Mt. Sohoton, 150–200m, 10–19 vii 1994, leg. T. Borromeo. Eastern Samar Province: ♂, San Rafael, 50–180m, 21–24 iv 1992, leg. T.

Borromeo jr; 3 ♂, ♀, San Rafael, 20–28 v 1993, leg. T. Borromeo; ♂, Taft, San Rafael, ix 2009, in coll. RJTV; ♀, Oras, 50–150m, 16–18 iv 1992, leg. T. Borromeo jr; 2 ♂, Oras, 16–17 iii 1994, leg. T. Borromeo; ♂, ♀, Oras, 21 iii 1994, leg. T. Borromeo; ♂, 2 ♀, Oras, 11–22 iv 1994, leg. T. Borromeo. Northern Samar: ♂, Las Navas, Brgy. San Isidro, 100–350m, 22–28 v 1997, leg. A. Buenafe”.

Etymology. A noun in the genitive case, named after the grandfather, Feliculo G. Torayno, of the first author.

Description of holotype male. Head: Labium pale, mandible bases pale with large dark mark. Labrum shiny black except for yellow distal third. Genae pale, pale colour continued to eye margin and narrowly along eye margin from ridge of frons to level of antennal sockets. Anteclypeus pale blue with dark central marking adjacent to postclypeus. Postclypeus black. Frons black, with distinct ridge, anterior face with paired transverse yellow streaks separated centrally. Vertex black with metallic reflection. Broad vertical pale streak on antennal socket, top of scape pale, pale vertical streak on pedicel, rest of antennae brown. Remainder of head dark metallic green. Distinct tubercle situated anteriorly beside eye margin in postocular area.

Thorax: Prothorax dark metallic green except for pale transverse streak on crest of anterior lobe and pale lower part of propleuron. Shelf of posterior pronotal lobe (as in Fig. 13 , which shows a paratype) short, with corners raised. Horns of posterior pronotal lobe long, rearward directed, abruptly bent outward at right angle apically, tapering to tip ( Fig. 13 ). Synthorax ( Fig. 35 ) with long process at ca 1/4 of length of dorsa carina, directed upward and slightly forward. Mesinfraepisternum metallic green with partial, irregular pale borders. Mesepisternum metallic green. Mesepimeron mostly metallic green with broad pale streak along interpleural suture, not reaching antealar carina ( Fig. 35 ). Metepisternum pale, large metallic green marking adjacent to antealar carina, separated from small dark mark spanning metapleural suture, metepimeron otherwise pale. Legs with coxa and trochanter pale, femur and tibia pale with dark spines, blackish streaks on extensor surfaces and dark markings around joint. Tarsi without denticle. Wings hyaline with black veins. Arc just off Ax2, Ac near to Ax2, petiolation ceases before Arc. R4 arising at or a little distal to subnodus; IR3 slightly more distal. 14 Px in Fw, 13 Px in Hw. Pt brown, almost rectangular, costal side a little shorter than subcostal side.

Abdomen: S1–2 metallic black dorsally, becoming pale lower on sides; S3–7 dorsally black, paler laterally with narrow pale basal ring. S8–10 black, paler lower laterally. Cerci pale, more than twice length of S10. Fork of upper branch at ca 1/3 length of upper branch ( Fig. 48 ). In lateral view upper branch shaped as in Fig. 60 , with small black internal tooth at tip, in dorsal view slightly hollowed interiorly toward tips. Spur ca 1/2 length of distance from S10 to fork in dorsal view, longer than wide and well visible in lateral view, tuft of dense setae at tip. Lower branch short, white, barely visible in lateral view with tip ca halfway from S10 to base of spur. Paraprocts typical for genus.

Measurements (mm): abdomen including cerci 39, Hw 23.

Female. Similar to male except for lateral portion of posterior pronotal lobe curved upward instead of out ( Fig. 25 ). S8–9 lateral pale marking more extensive with bluish tint, S9 with black restricted to dorsum.

Measurements (mm): abdomen 38, Hw 24.

Diagnosis. The very long, flat tubercle of the dorsal carina of the synthorax and the right angle outward turn of the apical part of the horns of the posterior pronotal lobe serve to separate this species from all others. S. dentifer from Dinagat and Siargao Islands also possess a long tubercle on the dorsum of the synthorax but lack the lower branch of the cercus and differ in the posterior lobe of the prothorax.

Figures

FIGURE 65. Distribution of S. braulitae (black circle), S. carmelae (black square) and S. feliculoi (black triangle).

FIGURES 8 – 15. Male posterior pronotal lobe, dorsal views: (8) S. braulitae Camiguin; (9) S. braulitae Homonhan; (10) S. bulba holotype; (11) S. circularis Tawi-Tawi; (12) S. dentifer Mindanao, Davao Oriental; (13) S. feliculoi paratype; (14) S. furcata; (15) S. zamboanga holotype.

FIGURES 19 – 26. Female posterior pronotal lobe, dorsal views: (19) S. bukid paratype; (20) S. bulba paratype; (21) S. carmelae paratype; (22) S. circularis Tawi-Tawi; (23) S. dentifer Dinagat; (24) S. dentifer Mindanao; (25) S. feliculoi paratype; (26) S. furcata.

FIGURES 35 – 40. Lateral views of male synthorax: (35) S. feliculoi holotype; (36) S. furcata; (37) S. hamis holotype; (38) S. janvantoli holotype; (39) S. zamboanga holotype; (40) Prothorax and part of synthorax of S. furcata male in oblique lateral view showing tubercle bearing clump of setae on tubercle on mesepisternum.

FIGURES 47 – 52. Lateral view of male anal appendages: (47) S. dentifer Mindanao; (48) S. feliculoi holotype; (49) S. furcata; (50) S. hamis holotype; (51) S. janvantoli paratype; (52) S. zamboanga paratype.

FIGURES 59 – 64. Dorsal view of male anal appendages: (59) S. dentifer Mindanao; (60) S. feliculoi holotype; (61) S. furcata; (62) S. hamis holotype; (63) S. janvantoli holotype; (64) S. zamboanga paratype.