Amphicnaeia fuscofasciata Wappes, Santos-Silva and Galileo, 2019

Wappes, James E., Santos-Silva, Antonio & Galileo, Maria Helena M., 2019, Synonymies, a new combination, and four new species in Amphicnaeia Bates (Coleoptera: Cerambycidae: Lamiinae), Insecta Mundi 703 (703), pp. 1-18 : 5-7

Amphicnaeia fuscofasciata Wappes, Santos-Silva and Galileo , new species

( Fig. 18–21 )

Description. Male. Integument mostly dark-brown; mouthparts yellowish-brown; distal area of labrum yellowish-brown; scape, pedicel, and antennomeres III–V reddish-brown, except dark brown apex of III–V; antennomeres VI–VIII reddish-brown basally, gradually and irregularly darker toward apex; antennomeres IX–XI with narrow reddish-brown basal ring, dark-brown on remaining surface; basal half of elytra with wide reddish-brown band, longitudinal on basal sixth, then curved inward and following longitudinally close to suture, and projected toward epipleural margin at same point of the curvature; femora mostly dark reddish-brown, more dark-brown in some irregular areas.

Head. Frons moderately coarsely and abundantly punctate; with yellowish-brown pubescence not obscuring integument, sparser in transverse area close to clypeus; with a few long, erect, brownish setae close to eyes. Vertex and area behind upper eye lobes with sculpturing as on frons; with light yellowish-brown pubescence nearly obscuring integument between antennal tubercles and upper eye lobes, following as pubescent band in each side of median groove, sparser close to superior area of upper eye lobes, denser, nearly obscuring integument close to lower eye lobes; remaining surface with brownish pubescence not obscuring integument; with a few long, erect brownish setae between antennal tubercles and close to upper eye lobes. Area behind lower eye lobes moderately coarsely, abundantly punctate (punctures coarser than behind upper eye lobes); with light yellowish-brown pubescence obscuring integument. Antennal tubercles coarsely, sparsely punctate basally, gradually finer, sparser toward smooth apex; pubescence as on frons. Genae finely, sparsely punctate except narrow apex smooth; with light yellowish-brown pubescence partially obscuring integument, except apex glabrous; with a few long, erect brownish setae. Postclypeus with light yellowish-brown pubescence in wide central area, partially obscuring integument, glabrous laterally. Labrum with sparse, long, erect yellowish setae. Distance between upper eye lobes 0.14 times length of scape; in frontal view, distance between lower eye lobes 0.83 times length of scape. Antennae 1.9 times elytral length, reaching elytral apex at distal third of antennomere VIII. Scape asperate dorsally, more distinctly so toward apex; with yellowish pubescence not obscuring integument; with long, erect, thick dark setae ventrally on distal third. Pedicel and antennomeres with long, erect dark setae ventrally, gradually shorter, sparser toward XI (interspersed with shorter, finer dark setae from V); dorsal surface of antennomeres with sparse, short, erect, brownish setae. Antennal formula (ratio) based on length of antennomere III: scape = 1.38; pedicel = 0.29; IV = 1.38; V = 1.33; VI = 1.33; VII = 1.33; VIII = 1.14; IX = 1.09; X = 0.90; XI = 1.00.

Thorax. Prothorax slightly wider than long; sides uniformly rounded, slightly narrowed posteriorly. Pronotum moderately coarsely, abundantly punctate; with dense light yellowish-brown pubescence laterally and in wide central band (somewhat wider centrally); remaining surface with light yellowish-brown pubescence not obscuring integument; with a few long, erect, thick brownish setae laterally on anterior and posterior area. Sides of prothorax with sculpturing as on pronotum, and pubescence as on sides of pronotum. Prosternum coarsely, moderately abundantly punctate; with yellowish-white pubescence not obscuring integument, slightly denser laterally. Prosternal process with sparse yellowish-white pubescence anteriorly, nearly glabrous posteriorly except a few long, erect, thick dark setae. Sides of ventral surface of meso- and metathorax with dense light yellowish-brown pubescence partially obscuring integument, and central area with yellowish-white pubescence not obscuring integument. Narrowest area of prosternal process about 4.5 times width of procoxal cavity. Narrowest area of mesoventral process about 3.5 times width of mesocoxal cavity. Scutellum with dense light yellowish-brown pubescence. Elytra. Coarsely, abundantly punctate on basal third, gradually sparser toward apex, especially from midlength; light area of anterior half with dense, light yellowish-brown pubescence, not obscuring punctures; with two, longitudinal, light yellowish-brown pubescent bands in anterior sixth, one from base to light area, between humerus and scutellum, another from scutellum to light area; with narrow, longitudinal light yellowish-brown pubescent band in sides of central third; with wide, longitudinal light yellowish-brown pubescent band on distal third, not reaching suture on its anterior area, bifurcated about its midlength, reaching suture near apex, then following laterally along epipleural margin, also only on distal third; remaining surface with brownish pubescence not obscuring integument; with long, erect, thick, dark setae throughout; apex nearly rounded. Legs. Femora with yellowish-white pubescence not obscuring integument (whiter depending on light intensity); with a few long, erect dark setae dorsally. Tibiae with light yellowish-white pubescence not obscuring integument on basal half, bristly, yellowish-brown on distal third; with sparse, long, erect dark setae.

Abdomen. Ventrites with moderately dense light yellowish-brown pubescence laterally, gradually sparser, yellowish-white toward center; distal margin of ventrite V slightly rounded.

Dimensions (mm). Total length 4.30; prothoracic length 0.90; anterior prothoracic width 0.85; posterior prothoracic width 0.80; maximum prothoracic width 0.95; humeral width 1.20; elytral length 2.95.

Type material. Holotype male from PANAMA, Panama: Cerro Azul ( Altos de Pacoras ), 4–10.I.1994, J.E.Wappes col. ( FSCA, formerly ACMT).

Remarks. Amphicnaeia fuscofasciata new species, differs from A. martinsi Galileo, 2015 (see photograph of the holotype at Bezark 2019) as follows: distance between upper eye lobes shorter than width of one lobe; central pubescence band of pronotum distinctly wider; anterior dense pubescence of the elytra not H-shaped. In A. martinsi , the distance between upper eye lobes is equal to width of one lobe, central pubescent band of pronotum narrow, and anterior dense pubescence of the elytra H-shaped. It differs from A. birai Galileo, 2015 (see photograph of the holotype at Bezark 2019) by the dense anterior pubescence of the elytra not forming narrow rows (in A. birai , forming three narrow longitudinal rows, interconnected between them by wide transverse band before middle of the elytra). Amphicnaeia fuscofasciata differs from A. flavescens Martins & Galileo, 1999 , by the central pubescent band of the pronotum distinctly wider (narrow in A. flavescens ), and basal half of the elytra without longitudinal pubescent band (present in A. flavescens ). The new species differs from A. lineata ( Fig. 27–28 ) by the upper eye lobes distinct wider (narrow in A. lineata ), and by the elytral pubescent pattern. Amphicnaeia fuscofasciata can be separated from A. antennata ( Fig. 15–17 ) by the upper eye lobes wider (narrower in A. antennata ), central pubescent band of the pronotum wider (narrow in A. antennata ), central area of prosternal process distinctly wider (very narrow in A. antennata ), and by the elytral pubescent pattern. Finally, it differs from A. bezarki new species by the upper eye lobes wider (narrow in A. bezarki ) pronotum not parallel-sided anteriorly (parallel-sided in A. bezarki , central pubescent band of the pronotum distinct wider (narrow in A. bezarki ) and elytral pubescent pattern.

Etymology. Named for the broad, dark fascia (resembling the batman shadow) that dominates the distal half of the elytra.

Figures

Figures 15–21. Amphicnaeia spp. 15–17) Amphicnaeia antennata, holotype male. 15) Dorsal habitus. 16) Ventral habitus. 17) Head, side view. 18–21) Amphicnaeia fuscofasciata, holotype male. 18) Lateral habitus. 19) Head, frontal view. 20) Dorsal habitus. 21) Ventral habitus.

Figures 22–28. Amphicnaeia spp. 22–26) Amphicnaeia rileyi. 22) Dorsal habitus, holotype male. 23) Ventral habitus, holotype male. 24) Lateral habitus, holotype male. 25) Head, frontal view, holotype male. 26) Dorsal habitus, paratype female. 27) Amphicnaeia affinis, holotype, dorsal habitus. 28) Amphicnaeia lineata, holotype, dorsal habitus.