Amara (Curtonotus) tumida Morawitz, 1862

Li, Yihang, Li, Haoyuan & Shi, Hongliang, 2024, Revision of the macropterous subgenus Curtonotus from east China, with the description of a new species (Carabidae, Zabrini, Amara), ZooKeys 1190, pp. 39-73 : 39

Amara (Curtonotus) tumida Morawitz, 1862

Amara (Leirus) tumida Morawitz, 1862: 258 (type locality: “Zagan-olui” [in Zabaykalsky Krai, Russia]; lectotype in ZRAS); Tschitschérine 1894: 390; Hieke 1990: 259 (lectotype designation).

Leirus tibialis Motschulsky, 1844: 343 (type locality: “Kamchatka”; holotype in ZMUM); junior secondary homonym of Amara (Amara) tibialis Paykull 1798; synonymized by Tschitschérine 1894: 390.

Amara (Curtonotus) tumida tunkunensis Hieke, 1990: 265 (type locality: "Quellgebiet des fl. Irkut" [= source of Irkut River, Buryatia, Russia]; holotype in MNHU).

Chinese common name.

膨胸暗步甲.

Diagnosis.

Small- to medium-sized species, BL = 9.0-11.0 mm; dorsum black, legs dark brown; head small, ~ 1/2 of pronotum maximum width, with one supraorbital seta. Pronotum cordate, widest near middle, densely punctate at base, sparsely punctate at mid-anterior region; lateral margins sinuate before posterior angles; posterior angles laterally protruded, acute or nearly rectangular. Elytra relatively long, widest near middle; lateral sides of abdominal sternites densely punctate. Male mesotibiae projection composed of two denticles; proximal denticle acutely pointed a little beyond middle; distal denticles smaller, present between the proximal one and tibial distal apex. Male genitalia with apical lamella slightly declined rightward, narrowed to apex; gonocoxite 2 of ovipositor elongated, length ~ 2.3 × as greatest width, apex narrow.

Comparison.

This species is most similar to A. (C.) shinanensis , which can only be distinguished from it by having longer apical lamella, more constricted pronotum base, and relatively longer body. Compared with A. (C.) gansuensis , A. (C.) tumida has more distinct mesotibiae denticles and narrower pronotum lateral groove; compared with A. (C.) goniodera , A. (C.) tumida has more sinuate lateral margins before posterior angles, longer gonocoxite and shorter apical lamella; compared with A. (C.) hyperborea , A. (C.) tumida has punctate abdominal sternites and more constricted pronotum base.

Distribution.

China (Heilongjiang, Inner Mongolia), Russia (east Siberia).

Remarks.

According to Hieke (1990), this species has shortened wings, to ~ 1/3 the length of elytra. We could not confirm it based on current material. If this is true, it will be an important feature to distinguish A. (C.) tumida from similar species, like A. (C.) shinanensis .