Allotrichoma (Allotrichoma) lasiocercum Cresson

Mathis, Wayne N. & Zatwarnicki, Tadeusz, 2012, Revision of New World Species of the Shore-fly Subgenus Allotrichoma Becker of the Genus Allotrichoma with Description of the Subgenus Neotrichoma (Diptera, Ephydridae, Hecamedini), ZooKeys 161, pp. 1-101 : 39-41

Allotrichoma (Allotrichoma) lasiocercum Cresson Figs 3338

Allotrichoma lasiocercum Cresson 1926: 251; 1948: 260 [synonymy with Allotrichoma simplex ]. Wirth 1965: 736 [Nearctic catalog]. Cole 1969: 397 [list]. Mathis and Zatwarnicki 1995: 151 [world catalog].

Description.

This species is distinguished from congeners by the following combination of characters: Small to moderately small shore flies, body length 1.55-2.15 mm. Head: Medial facial carina above facial prominence shallow; labella broad, fleshy, shorter than mediproboscis; clypeus microtomentose, usually gray.

Thorax: Presutural supra-alar seta present. Pleural area lacking stripes. Wing with costal vein ratio 0.36-0.38; M vein ratio 0.37-0.40. Legs blackish brown to black; midtibia bearing numerous, erect setae along dorsal surface.

Abdomen: Male 5th sternal flap in lateral view (Fig. 37) pedunculate, relatively narrow at base, becoming wider apically, bearing numerous setulae along truncate apex; 5th medial process in lateral view (Fig. 37) elongate, bar-like, bearing 4-5 apical setulae. Male terminalia (Figs 33-36): Epandrium in posterior view (Fig. 33) ovately rounded on dorsal half; epandrium in lateral view (Fig. 34) with anterior and posterior margins sinuous, relatively wide dorsally; cercus in posterior view (Fig. 33) broadly pointed dorsally, gradually broadened ventrally then abruptly curved or arched laterally and ventrally as a slightly tapered, elongate, arched process that bears numerous moderately long setulae apically; cercus in lateral view (Fig. 34) shallowly arched, wider dorsally, ventral half nearly parallel sided, apex broadened and bearing moderately long setulae; surstyli (ventral extensions of epandrium) in posterior view as parallel, narrow processes that bear setulae on apical portion; surstylus in lateral view narrowed and nearly parallel sided, apical portion bearing setulae anteriorly and apically, with a short, apical process that bears a longer, apical setula; aedeagus in ventral view (Fig. 35) elongate, narrowly ovate, tapered apically, in lateral view (Fig. 36) elongate, somewhat lunate; phallapodeme in lateral view (Fig. 36) triangular, keel shorter than length, pointed; gonite in ventral view (Fig. 35) bar-like, slightly wider basally, apex with short, lateral process, apex bearing setula, in lateral view (Fig. 36) wide basally, gently but distinctly arched, tapered to apical point.

Type material.

The holotype male is labeled "Berkeley Hills[,] Alameda Co.[,] IV,11,'08 [11 Apr 1908], Cal[ifornia]/♂/241/TYPE Allotrichoma, LASIOCERCUM, E.T. Cresson, Jr [red; species and generic names handwritten]/Loaned by A.N.S.P." The holotype male is double mounted (minuten in a small rectangular card), is in good condition (abdomen removed, dissected, stored in glycerin in an attached microvial), and is deposited in the ANSP (6319). Cresson also listed two female paratopotypes (ANSP).

Type locality.

United States. California. Alameda: Berkeley Hills (37°53'N, 122°14'W).

Other specimens examined from the New World.

UNITED STATES. CALIFORNIA. Inyo: Tecopa Hot Spring (35°52.7'N, 116°13.9'W), 12 Apr 2003, W. N. Mathis, T. Zatwarnicki (1♀; USNM). Marin: Novato, San Jose Creek (38°06.5'N, 122°34.2'W; 45 m), 10 Jul 1971, M. M. and P. H. Arnaud, Jr. (1♂; CAS). Mendocino: Russian River near Hopland (38°58.4'N, 123°07'W), 4 May 1968, W. J. Turner (1♂; WSU). Mono: Mammoth lakes (37°36.3'N, 119°0.7'W), 29 Jul 1940, L. C. Kulters (1♂; KU); Mono Lake (38°01'N, 119°0.1'W), 16 Aug 1958 (1♂; USNM). Riverside: Martinez (33°33.8'N, 116°09.2'W), 11 Jul 1917, J. M. Aldrich (1♂; USNM). San Bernardino: Saratoga Spring (35°40.9'N, 116°25.3'W; 100 m), 31 Mar 2005, D. and W. N. Mathis (2♂, 7♀; USNM). Santa Clara: San Jose (37°20.4'N, 121°53.7'W), 8 Sep 1949, L. W. Quate (1♂, 1♀; USNM); San Jose, Guadalupe Creek (37°14.8'N, 121°52.3'W), 14 Jun 1966, R. E. Orth (1♂; UCR). San Diego: Carlsbad (33°09.5'N, 117°21'W), 1 Jun 1954, J. C. Hall (1♂; USNM).

Distribution.

(Fig. 38) Nearctic: United States (California).

Remarks.

Cresson (1929: 176) suggested that a male from Vienna, Austria (Mik), was conspecific with this species. Cresson was probably mistaken and may have confused Allotrichoma bezzii with this species. A second Palearctic species, Allotrichoma picenum Canzoneri and Rampini (1990: 44; Italy), is similar to Allotrichoma lasiocercum but has shorter cerci.