Allococalodes madidus, Maddison, Wayne P., 2009

Maddison, Wayne P., 2009, New cocalodine jumping spiders from Papua New Guinea (Araneae: Salticidae: Cocalodinae), Zootaxa 2021, pp. 1-22 : 5-7

Allococalodes madidus new species

( Figs 10–19 )

Type material. Holotype male in UBC – SEM with labels: " PAPUA NEW GUINEA: Southern Highlands Province: Umgé. S 5.3054 E 142.5098 and S 5.3059 E 142.5111. 1400 m a.s.l. 16–17 July 2008. W. Maddison & Aislan Tama Wanakipa Indiaf. WPM#08–014", " Allococalodes madidus Maddison , Holotype " " UBC ––SEM AR 00020". Paratypes: 3 females, 1 male, 9 juveniles with same collecting data.

Etymology. The Latin madidus means wet or moist, referring to the habitat of mossy vines and trunks along a forest stream.

Diagnosis. Tibial apophyses of male palp much smaller than in A. alticeps Wanless and A. cornutus Wanless , not projecting retrolaterally as in those species.

Description. Male (holotype). Carapace length 2.5; abdomen length 2.2. Chelicerae elongate, projecting forward, in life held together ( Fig. 13 ). Four promarginal and two retromarginal teeth. Intercheliceral sclerite elongated into a horn as in other Allococalodes and Cocalodes . Palpus ( Figs 16–17 ) with embolus short and accompanied by a conductor and median apophysis, as in the other two species of Allococalodes ( Wanless 1982) . Median apophysis flat, leaf-like. Tibia of first leg with three pairs of ventral macrosetae; first metatarsus with three pairs. Carapace high, with posterior eyes on tubercles and area of fovea concave. PME large. Colour ( Figs 10, 11, 13 ): carapace and abdomen dark brown except paler longitudinal stripe on thoracic slope and dorsally on abdomen. Recumbent golden-orange setae concentrated near eyes and over heart; white setae over thoracic stripe; black elsewhere. Palpi pale olive yellow except cymbium red-brown. Legs somewhat annulate olive and black.

Female (paratype, UBC–SEM AR00020). Carapace length 2.9; abdomen length 3.4. Chelicerae: three promarginal and three retromarginal teeth. Palp with tarsal claw. Tibia of first leg with three pairs of ventral macrosetae; first metatarsus with 3 pairs. Epigynum ( Fig. 18 ) with two small posterior openings. Carapace high, with posterior eyes on tubercles and area of fovea concave. PME large. Colour ( Figs 12, 14, 15 ): as in male.

Natural history. All specimens were found on moss-covered surfaces within a few metres of a shaded stream in the forest at mid elevation. Most were on moss-covered trunks of small trees and vines ( Fig. 87 ).

Figures

FIGURES 10 – 15. Allococalodes madidus, new species. 10 – 11 Holotype male. 12, 14, 15 Paratype female. 13 Paratype male.

FIGURES 16 – 19. Allococalodes madidus, new species. 16 Palp of holotype male, ventral view. 17 Palp of holotype male, retrolateral view. 18 Epigynum, ventral view. 19 Cleared epigynum, dorsal view. Scale bars 0.1 mm.

FIGURES 87 – 92. Habitats of cocalodines. Microhabitat indicated by arrows. 87 Habitat of Allococalodes madidus, on mosses on small tree trunks along stream (Umgé, S 5.3054 E 142.5098); approximate width of field 1.5 m. 88 Habitat of Yamangalea frewana, on tree branches and trunks (Mt. Gahavisuka, S 6.015 E 145.412); approximate diameter of branch indicated 5 cm. Most were found on larger, vertical tree trunks. 89 Habitat of Tabuina varirata, on conifer in clearing (Varirata National Park, S 9.436 E 147.364); approximate width of field 0.7 m. 90 Habitat of Cucudeta zabkai, in leaf litter at base of Pandanus (near Tualapa, S 5.291 E 142.500); approximate width of field 1 m. 91 Habitat of Cucudeta uzet, in leaf litter at base of Pandanus (Umgé, S 5.304 E 142.512); approximate width of field 1 m. 92 Habitat of Cucudeta gahavisuka, in leaf litter on terraces on slope in forest (Mt. Gahavisuka, S 6.01451 E 145.41171); approximate width of field 1.5 m.