Agyneta fratrella ( Chamberlin 1919 )

Dupérré, Nadine, 2013, Taxonomic revision of the spider genera Agyneta and Tennesseellum (Araneae, Linyphiidae) of North America north of Mexico with a study of the embolic division within Micronetinae sensu Saaristo & Tanasevitch 1996, Zootaxa 3674 (1), pp. 1-189 : 80-83

Agyneta fratrella ( Chamberlin 1919)

Figs 238–245 , map 16.

Grammonota fratrella Chamberlin 1919: 250 , pl. 18, f. 3. (Description Ƥ).

Microneta fratrella Chamberlin & Ivie 1933: 35 , pl. 6, f. 61, pl. 9, f. 93–95, pl. 10, f. 101–102. (Ƥ, description 3). Meioneta fratrella Lowrie & Gertsch 1955: 5 . (Transferred Ƥ from Microneta ).

Agyneta fratrella Buckle et al. 2001: 100 . (Transferred from Meioneta ).

Type material: Grammonota fratrella Chamberlin 1919 , Ƥ HOLOTYPE from Utah, Uinta Mts. 8500ft. R.V. Chamberlin Coll. (unique number 443–RVC). MCZ, EXAMINED.

Diagnosis: Males are diagnosed from all species by the scimitar-shaped lamella characteristica ( Fig. 238 ). From other species in the group by their longer, pointed cymbial ventral tubercle ( Fig. 239 ), shorter and rounded in A. danielbelangeri and A. pistrix ( Figs 247, 251 ). Females are diagnosed from all Agyneta by the narrow smileshaped lateral lobes ( Fig. 242 ).

Description: Male: Total length 1.71; carapace length 0.82, width 0.66.

CEPHALOTHORAX: Carapace dark orange-brown, shiny, finely reticulate; margin, radiating lines and pars cephalica lightly suffused with dark gray; trident mark present. Sternum orange-brown strongly suffused with dark gray. Clypeus height 2. Chelicerae dark orange-brown, excavated, ~ 16 seta-tipped tubercles; promargin one tooh, retromargin four tiny denticles on margin, two tiny denticles in between margins; both margins with large projections near base of fangs. Cheliceral stridulatory organ visible, ~ 38 striae, well spaced slowly getting closer basally. ABDOMEN: Uniformly medium gray. LEGS: Orange-brown, femur darker; leg I total length: 3.32; leg III total length: 2.34; Tm I: 0.27, Tm IV: absent. GENITALIA: Palpal retrolateral tibial apophysis well sclerotized, with small pointed tip; dorsal tibial apophysis with rounded tip, smooth, not strongly sclerotized; two retrolateral and one dorsal trichobothria ( Fig. 238 ). Cymbium triangular; glabrous depression present ( Fig. 238 ); dorsal cymbial tubercle anvil-shaped and smooth; ventral turbercle long, acute and smooth; prolateral notch deep ( Fig. 239 ). Paracymbium apically constricted, apical pocket long, anterior pocket short, deep and curved making a medium cover, posterior pocket absent ( Fig. 238 ). Embolus tip pointed with apical prong; ventro-retrolaterally with rows of spines; basally excavated with basal flange; Fickert’s gland elongated set basally; ventral lamella pointed, transparent; thumb elongated, reaching beyond the embolus proper, with well sclerotized tip ( Fig. 240 ). Embolus proper set basally ( Fig. 240 ). Anterior terminal apophysis long with long protrusions; posterior terminal apophysis small and twisted with large basal prong; lamella characteristica with bifid pointed tip ( Fig. 241 ).

Female: Total length 1.94; carapace length 0.73, width 0.59.

MAP. 16. Localities of Agyneta fratrella ( Chamberlin 1919) .

CEPHALOTHORAX: Same coloration as in male. Chelicerae promargin four teeth, retromargin four denticles. Cheliceral stridulatory organ ~30 striae, closely spaced. ABDOMEN: Same as male. LEGS: Same as male; palpal tarsal claw absent, leg I total length: 2.86; leg III total length: 2.06; Tm I: 0.23, Tm IV: absent. GENITALIA: Epigynum with wide proximal part of scape, sides angular; epigynal slits long; pit hook depression average ( Fig. 242 ); lateral lobes long, stretcher small; pit deep ( Fig. 243 ). Median part of scape very narrow ( Fig. 244 ); genital pores situated in median part of scape ( Fig. 245 ). Internal genitalia with large, oval receptacula set horizontally ( Fig. 243 ).

Other material examined: USA: Colorado: 11km S Ouray, 2956m, 1Ƥ ( AMNH); Idaho Springs, 24.viii.1935, 8Ƥ, W. Ivie ( AMNH); Upper Spring Creek, near Monarch Pass, ix.1937, 1 Ƥ, R. Chamberlin ( AMNH); Westcreek, 05.viii.1941, 1Ƥ, S., D. Mulaik ( AMNH). Idaho: 6km N McCall, 18.x.1944, 531Ƥ, W. Ivie ( AMNH); Lake Forks, 05.vii.1943, 2Ƥ, W. Ivie ( AMNH); Payette Lake, southwest shore, 04.vii.1943, 23, W. Ivie ( AMNH). New Mexico: Cimarron Canyon, W of Ute Park, 06.x.1965, J., W. Ivie ( AMNH); S-21, 10.xi.1949, 1Ƥ, C. Hoff ( AMNH). Utah: Brighton, 2651m, 20.xi.1939, 2Ƥ, S. Mulaik ( AMNH); Raft Mountains, Clear Creek, 04.ix.1943, 2Ƥ, R. Chamberlin ( AMNH); Uinta Mountains, Cobble Rest camp, 24.ix.1932, 1Ƥ ( AMNH); Uinta Mountains, Cobble Rest camp, North fork of Provo River, 24.ix.1932, 11Ƥ ( AMNH); Uinta Mountains, Lost Lake, 19.vi.1941, 1Ƥ, R. Chamberlin ( AMNH); Uinta Mountains, Mirror Lake, 22.ix.1932, 1Ƥ, 18.viii.1942, 1Ƥ, R. Chamberlin, W. Ivie ( AMNH); Vasquez, 24.viii.1935, 1Ƥ, W. Ivie ( AMNH). Washington: Bonanza Camp [47.386N, 120.599W] 914m, 10.viii.1980, moss, litter by stream, 1Ƥ, D. Carroll ( UWBM); Colockum Creek, north fork [47.24N, 120.296W] 1499m, rotten stump and riparian Salix litter, 4Ƥ, R. Crawford ( UWBM); Kamiak Butte [46.868N, 117.156W] 911–926m, 04.vi–26. viii.1994, 5 Ƥ, 05.vii–01. xi.1995, 1 Ƥ, pitfalls in forest, J. Bergdhal ( UWBM); junction of Deep and Careter Creeks, 21.vii.1935, 1Ƥ, R. Chamberlin ( AMNH); Lasal Mountains, 2438m, 29.x.1931, 4Ƥ, W. Ivie ( AMNH); Logan Canyon, 1Ƥ, R. Chamberlin ( AMNH); Lowries Canyon, 11.viii.1935, 1Ƥ, R. Chamberlin ( AMNH); Rabbit Creek [48.539N, 118.605W] 1066m, 18.vi.1989, alder litter, R. Crawford ( UWBM); Salmon Creek [48.503N, 119.725W] 560m, 14.vi.1984, sifting Populus litter, R. Crawford ( UWBM). Wyoming: Brooks Lake Falls, 12.viii.1941, 1Ƥ, S. Mulaik ( AMNH).

Distribution: Northwestern USA, south to Utah, Colorado and New Mexico.

Figures

FIGURES 238 – 245. Agyneta fratrella (Chamberlin 1919). 238 – 241, Male. 242 – 245, Female. 238. Palp, retrolateral view. 239. Palp, prolateral view (arrow points to embolus basal excavation). 240. Embolus, prolateral view. 241. Radical division, retrolateral view. 242. Epigynum, ventral view. 243. Internal genitalia, ventral view. 244. Internal genitalia, dorsal view. 245. Internal genitalia, lateral view.

FIGURES 246 – 253. 246 – 249. Agyneta danielbelangeri n. sp. 246 – 249, Male. 246. Palp, retrolateral view (arrow points to constriction of the apical part of paracymbiun). 247. Palp, prolateral view. 248. Embolus, prolateral view. 249. Radical division, retrolateral view. 250 – 253. Agyneta pistrix n. sp. 250 – 253, Male. 250. Palp, retrolateral view. 251. Palp, prolateral view. 252. Embolus, prolateral view. 253. Radical division, retrolateral view.