Aeshna zlatkokvaceki, Prokop & Pecharová & Nel, 2016

Prokop, Jakub, Pecharová, Martina & Nel, André, 2016, New Cenozoic dragonflies from the Most Basin and Středohoří Complex volcanic area (Czech Republic, Germany), Journal of Natural History 50, pp. 2311-2326 : 2313-2314

Aeshna zlatkokvaceki sp. nov.

( Figure 2 (a, b))

Etymology

The epithet honours Prof. Zlatko Kvaček (Charles University, Praha), a world-famous palaeobotanist who systematically evaluated plant macrofossils from this locality.

Holotype

SMMG CsT 1091 (a nearly complete fore wing preserved in salmon-pink baked clay – porcelanite), collection of Senckenberg Naturhistorische Sammlungen Dresden , Saxony, Germany.

Age and outcrop

Early Miocene (Burdigalian), Most Basin, Most Formation, Libkovice Member, Želénky near Duchcov ( Figure 1 ), Czech Republic (see Kvaček and Hurník 2000).

Description

A nearly complete fore wing, with only median area and extreme apex missing; whole wing surface apparently hyaline; wing c. 55 mm long (assumed from fragment) and 11.6 mm wide; distance from estimate base to nodus c. 26.6 mm; distance from nodus to wing apex, c. 28.2 mm; nodus nearly midway between base and apex; distance from nodus to pterostigma 16.3 mm; distance from pterostigma to apex about 7.5 mm; pterostigma rather long, 4.4 mm long and 0.9 mm wide, covering approximately three cells and a half; pterostigmal brace slightly obliquely aligned with proximal side of pterostigma; 11 postnodal cross-veins, not well aligned with 13 visible subpostnodal cross-veins; 19 visible antenodal crossveins of first row between C and ScP not aligned with the 17 visible corresponding antenodal cross-veins of second row between ScP and RA; 13 secondary antenodal cross-veins between Ax2 and nodus; four antenodal crossveins between Ax2 and Ax1; Ax1 4.6 mm and Ax2 10.9 mm from wing base; hypertriangle crossed by four cross-veins; median space not preserved; submedian space crossed by five cross-veins, subdiscoidal triangle three-celled; discoidal triangle elongate and divided into six smaller cells, its costal side being 8.1 mm long, distal side 6.9 mm long and proximal side 3.7 mm long; width of postdiscoidal area just behind discoidal triangle 3.4 mm, width along posterior wing margin 7.2 mm; three rows of cells in postdiscoidal area just distal of discoidal triangle; a short convex supplementary sector (trigonal planate) in postdiscoidal area, aligned with concave Mspl; Mspl well defined, nearly straight in its basal half but slightly curved in distal part; three rows of cells between Mspl and MP and three rows between Mspl and MA; two rows of cells and a short zigzagged supplementary vein in distal part of area between MA and RP3/4; bulge in distal part of MA (‘ aeshnid bulla ’) weak but present; four preserved Bq cross-veins; oblique vein ‘ O ’ one cell distal of base of RP2; Rspl well defined and nearly straight; area between Rspl and IR2 wide, with four rows of cells in its widest part; space between IR2 and Rspl basally divided by oblique intercalary veinlets; IR2 smoothly curved distally and asymmetrically forked just basal of pterostigma, with three rows of cells between its branches; RP2 strongly curved posteriorly opposite proximal side of pterostigma; one row of cells between RP2 and anterior branch of IR2; IR1 present but zigzagged and short, about 8.7 mm long, beginning just below distal side of pterostigma; one row of cells between MP and CuAa in basal parts but five rows along posterior wing margin; CuAa with circa five posterior branches directed towards posterior wing margin; cubitoanal area broad with six rows of cells below CuAa.

Figures

Figure 1. Map of Cenozoic dragonfly localities in the Krušné hory piedmont basins and volcanic areas. Abbreviations: I, Cheb and Sokolov basins; II, Most Basin; III, České středohoří Mts. and Doupovské hory Mts.; 1, Želénky near Duchcov; 2, Bílina mine; 3, Kundratice near Litoměřice; 4, Seifhennersdorf; 5, Libouš mine; 6, Jehličná near Sokolov; 7, Marie Majerová mine near Sokolov; 8, Pochlovice near Kynšperk n. Ohří.

Figure 2. Aeshna zlatkokvaceki sp. nov. (Aeshnidae) (A) Photograph of holotype specimen SMMG CsT 1091 (Senckenberg Naturhistorische Sammlungen Dresden coll., Germany), imprint only; (B) line drawing of fore wing. Scale bars represent 5 mm.