Phalangopsidae crickets from the Indian Region (Orthoptera, Grylloidea), with the descriptions of new taxa, diagnoses for genera, and a key to Indian genera Desutter-Grandcolas, Laure Jaiswara, Ranjana Zootaxa 2012 3444 1 39 8NKP3 [253,450,1066,1092] Insecta Phalangopsidae GBIF,CoL Animalia Orthoptera 3 4 Arthropoda family    1. Both males and females with long FWs covering the whole abdomen. TIII with three pairs of subapical spurs (  Meloimorpha), or less (  Cacoplistes).................................................................................... 2  - Males and females either apterous, or with short, more or less reduced FWs. TIII with four outer subapical spurs, and three (  Paragryllodes) or four inner subapical spurs................................................................ 3   2. Large species, with strong body and legs. Pronotum concave dorsally, its margins deeply carinated. TIII thick with strong latero-dorsal margins; subapical spurs very short; apical spurs widely set apart from each other on both inner and outer sides.....................................................................  CacoplistesBrunnervon Wattenwyl, 1873  - Small and thin species, with thin legs. Pronotum margins not carinated, but dorsal disc saddle-like. TIII without lateral carina, quadrangulate in section; subapical spurs long and clearly articulated; apical spurs close to each other on each tibia side...................................................................................  Meloimorpha Walker, 1870   3. TIII with three inner subapical spurs. Fastigium very narrow, much more narrow than the scape, and furrowed. Male subgenital plate elongate; FWs short; stridulum: harp crossed by many veins, mirror not well defined (Fig. 41 in Chopard 1928). Male genitalia typical for the genus, with a short, transverse pseudepiphallic sclerite and long ectophallic dorsal valves (Desutter- Grandcolas 1999). Female winged, FWs with projecting veins (Fig. 40 in Chopard 1928).............................................................................................................  Paragryllodes Karny, 1909  - TIII with four inner subapical spurs. Fastigium wider and not furrowed. Male apterous or winged; when the stridulum is present, harp crossed by no more than two or three veins; mirror present, reticulate or not. Male genitalia without ectophallic dorsal valves. Females apterous when known (except in  Luzaropsis)................................................... 4   4. TIII apical spurs: median spurs the longest on both inner and outer sides; on outer side, the median either greatly (  Arachnomimus,  Zacla?, Fig. 1J) or slightly (  Kempiola, Fig. 3C) longer than the dorsal spur................................... 5  - TIII apical spurs different............................................................................... 7   5. Both sexes apterous. TI without tympanum. Male genitalia: median process resembling those of  Zacla(cf. infra). Female genitalia long and horse-foot shaped, with a thinner basis and a wider, split apex, and with a plicated, basal membrane ( Fig. 1G, H).........................................................................  ArachnomimusSaussure, 1897  - Male FWs more or less reduced in length, but with a complete stridulum; female apterous. TI with an inner tympanum (except in  K. flavipunctatusDesutter-Grandcolas  n. sp.)............................................................ 6   6. TIII serrulation very sparse (except in  K. flavipunctatusDesutter-Grandcolas  n. sp.); TIII median and dorsal outer apical spurs subequal, the median only slightly longer than the dorsal one ( Fig. 3C). Male stridulum reticulated. Male genitalia: pseudepiphallus with a long and narrow median process, and elongate lateral lobes ( Figs 3F, 5B). Female copulatory papilla elongate and very flat, entirely sclerotized ( Fig. 3D, E, G–I).................................  Kempiola Uvarov, 1940  - TIII serrulation strong; TIII median outer apical spur clearly longer than the dorsal one. Male stridulum not reticulated (Fig.  166 in Chopard 1969). Male genitalia with a short and straight median lobe, slightly shorter than lateral sclerites (Fig. 4.1– 3 in Gorochov 2003a). Female genitalia unknown.............................................  ZaclaGorochov, 2003  7. TIII apical spurs: dorsal spur the longest on outer side, and dorsal spur the longest on inner side (except in P. (?) chopardiDesutter-Grandcolas  n. sp., in which the median is slightly longer than the dorsal). TI without a tympanum. Males apterous or with very short FWs. Females apterous, when known........................................................ 8  - TIII apical spurs: median spur the longest on outer side, and dorsal spur slightly longer on inner side. TI with an inner tympanum, or two tympana. Both sexes winged................  Larandopsis Chopard, 1924,  Luzaropsis Chopard, 1925   8. TIII not serrulated. Eyes reduced, as long as the scape. Male first abdominal tergite very large and produced above the other tergites in the shape of a triangular shield. Female unknown.............................  AspidogryllusChopard, 1933  - TIII serrulated. Eyes not as reduced. Male first abdominal tergite not modified...................................... 9   9. Pronotum distinctly transverse ( Figs 7C, G, 10E). Fastigium much wider than the scape ( Figs 7G, 10E)................. 10  - Pronotum less transverse ( Fig. 10A). Fastigium slightly more narrow than the scape ( Fig. 10B). Male unknown. Female copulatory papilla small and flat, flap-like ( Fig. 9O).........................  SpeluncasinaDesutter-Grandcolas n. gen.   10. TIII subapical spurs: outers much longer than the inners. Male apterous or with flap-like, largely separate FWs; lateral parts of supra anal plate elongate. Male genitalia resembling those of  Phalangopsina, but small, rounded and compact ( Fig. 12); pseudepiphallic sclerite median lobe flat; pseudepiphallic parameres having the shape of acute hooks ( Fig. 12B, C). Female ovipositor shorter than FIII; copulatory papilla flat, concave, sclerotized only distally ( Fig. 11H–J)...........................................................................................  OpiliosinaDesutter-Grandcolas n. gen.  - TIII subapical spurs: outers only slightly longer than the inners. Male FWs as long as tergite 1, slightly overlapping; supra anal plate simple. Male genitalia: pseudepiphallic sclerite with a wide median part, more or less reversed overhead, and two long lateral sclerites; pseudepiphallic parameres very large, concave, making a kind of wide wide forceps ( Fig. 6D–F). Female ovipositor longer than FIII; copulatory papilla large, sclerotized and elongate ( Figs 6G–H, 8F, G), with a longer and thinner distal part (shorter in P. (?) chopardiDesutter-Grandcolas  n. sp., see Fig. 8K, L)..........  PhalangopsinaChopard, 1933