Revision of the Oriental genus Horniella Raffray (Coleoptera, Staphylinidae, Pselaphinae) Yin, Zi-Wei Li, Li-Zhen Zootaxa 2014 2014-08-12 3850 1 1 83 6MDCM Yin and Li Yin and Li [195,615,1546,1572] Insecta Staphylinidae Horniella Animalia Coleoptera 41 42 Arthropoda species asymmetrica sp. nov.   Figs 24A, 25, 48H; Map 4    Typematerial(2 ♂♂, 5 ♀♀).  Holotype, ♂, labeled ‘ THAILAND, Khao Yai Nat. Park,  26.xi–3.xii.1985, 750– 850 m, Burckhardt-Löbl/ Holotype [red], ♂,  Horniella asymmetrica  sp. n., det. Yin& Li, 2014, MHNG’. Paratypes:  1 ♂, 3 ♀♀, same data as the holotype;  1 ♀, labeled ‘ THAILAND: Khao Yai Nat. Park,  4.–5.v.1978, P. Schwendinger’;  1 ♀, labeled ‘ THAILAND: Chanthaburi, Khao Sabap Nat. Park,  150–300 m,  23–24.xi.1985, Burckhardt-Löbl’. All paratypesare housed in MHNG, and each bears a yellow typelabel similar to that of the holotypeexcept ‘ PARATYPE♂ (or ♀)’.   Description.Male ( Fig. 24A). Length 3.03–3.15 mm. Head about as long as wide, HL 0.65–0.67 mm, HW 0.63–0.64 mm; anterolateral genal projections ( Fig. 25C) distinct, anterior margins deeply and roundly concave; median sulcus between antennal tubercles short; scapes ( Fig. 25B) with angulate expansion at basolateral margins; clubs ( Fig. 25A) loosely formed by apical three moderately enlarged antennomeres; venter with pair of thick lateral spines ( Fig. 25D). Maxillary palpomeres II stout, widest near middle. Each eye composed of about 48 facets. Pronotum slightly longer than wide, PL 0.63–0.65 mm, PW 0.60–0.61 mm. Elytra wider than long, EL 0.81–0.87 mm, EW 1.26–1.29 mm; discal striae reaching apical 3/4 of elytral length. Protrochanters and profemora ( Fig. 25E) each with one long ventral spine, protibiae ( Fig. 25F) each with small triangular preapical denticle; mesotrochanters ( Fig. 25G) with large, apically truncate ventral projection, mesofemora simple, mesotibiae ( Fig. 25H) with small bluntly triangular preapical denticle; tarsomeres II normal, not extending to beneath tarsomeres III. Abdomen large, AL 0.94–0.96 mm, AW 1.27–1.28 mm, tergite IV (first visible tergite) with median carina extending to half tergal length, lacking lateral discal carinae, tergite V lacking median carina. Sternite IX ( Fig. 25I) elongate, with well-sclerotized apical half, and membranous basal half. AeL 0.63 mm; aedeagus ( Figs 25J–L) with strongly asymmetric median lobe conspicuously cured toward right at apex; endophallus composed of three sclerites: one broad sclerite with curved branch at middle, one elongate, curved sclerite overlapping with broad one at base, and one much shorter sclerite beside elongate one. Female. Similar to male in general appearance; each eye composed of about 25–30 facets; profemora each with two ventral spines near base, apices of pro- and mesotibiae lacking denticle. BL 2.84–3.09 mm, HL 0.64–0.67 mm, HW 0.55–0.58 mm, PL 0.59–0.61 mm, PW 0.58–0.61 mm, EL 0.70–0.78 mm, EW 1.17–1.26, AL 0.91–1.03 mm, AW 1.19–1.31 mm. Genital complex ( Fig. 48H) weakly sclerotized, with transverse apical sclerite.   Differential diagnosis.This distinctive species is placed as a member of the  H. burckhardtigroup. The relatively large compound eyes, and the large, apically truncate ventral projection of the mesotrochanters are unique, which when combined with the strongly asymmetric form of the aedeagal median lobe lead to a ready separation of  H. asymmetricafrom all other congeners of the group.   Distribution. Thailand: Prachin Buri, Chanthaburi ( Map 4).   Collection notes.Adults were collected from leaf litter samples by sifting and use of Winkler-Moczarski extractors.   Etymology.The specific name ‘  asymmetrica’ refers to the strongly asymmetric aedeagal median lobe of the new species. 3699171302 1985-11-26 1985-12-03 1985-11-26 MHNG Burckhardt-Lobl Thailand Yin & Li 0 Khao Yai Nat. Park 41 42 1 1 holotype 3699171305 1985-11-26 1985-12-03 1985-11-26 MHNG Burckhardt-Lobl Thailand Yin & Li 0 Khao Yai Nat. Park 41 42 4 3 1 paratype 3699171317 1978-05-04 1978-05-05 1978-05-04 MHNG P. Schwendinger' Thailand Khao Yai Nat. Park 41 42 1 1 paratype 3699171309 1985-11-23 1985-11-24 1985-11-23 MHNG Burckhardt & Lobl Thailand 225 Khao Sabap Nat. Park Chanthaburi 41 42 1 1 paratype