A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae) Mbo, Zingisile Haddad, Charles R. Zootaxa 2019 2019-04-10 4582 1 1 62 37LKT Mbo & Haddad, 2019 Mbo & Haddad 2019 [151,413,1015,1041] Arachnida Gallieniellidae Drassodella GBIF Animalia Araneae 23 24 Arthropoda species maculata sp. nov.   Figs 63, 98, 123, 124   Etymology.Taken from the Latin maculatus, meaning spotted, referring to the white spots on the dorsal surface of the abdomen.   Diagnosis.  Drassodella maculata  sp. nov.females are easily recognised from other species in the  D. melanagroup by very shallow ( Fig. 63) rather than deep anterior hood (e.g. Figs 60–62), and the ST II that are much smaller than ST I. Male unknown.   Description. Female ( Holotype, Asante Sana Game Reserve, NCA 2012/4162).Measurements: CL 2.18, CW 1.70, AL 3.16, AW 1.78, TL 5.64 (5.25–6.30), FL 0.19, SL 1.25, SW 1.01, CH0.12, AME-AME 0.05, AME- ALE 0.02, ALE-ALE 0.20, PME-PME 0.08, PME-PLE 0.10, PLE-PLE 0.98, PERW 0.46, MOQAW 0.14, MOQPW 0.19, MOQL 0.07. Length of leg segments: I 1.32 + 0.76 + 1.14 + 1.02 + 0.88 = 5.12, II 1.36 + 0.66 + 0.92 + 1.00 + 0.76 = 4.70, III 1.18 + 0.66 + 0.82 + 1.06 + 0.70 = 4.42, IV 1.92 + 0.58 + 1.76 + 1.92 + 0.64 = 6.82. General appearance as in Fig. 98. Carapace orange-brown, with black radiating mottling, more intense towards posterior of cephalic region; few white setae across surface, white thin line at lateral edges of feathery setae, median white line absent, thin white lines of few white setae and black erect setae diverging behind fovea towards posterior margin. Clypeus height equal to AME diameter; AME smaller than ALE; AME separated by distance slightly smaller than their diameter, AME separated from AME by distance equal to 0.33 AME diameter; PME smaller than PLE; PME separated by distance equal to 1.33 their diameter, PME separated from PLE by distance equal to 0.17 PME diameter. Chelicerae orange; promargin with two teeth closely spaced; retromargin with one tooth. Endites orange. Labium orange anteriorly, dark-brown posteriorly, with very dark corners. Sternum orange; setae concentrated around margin, few in centre. Legs brown proximally, orange distally. Leg spination: femora: I do 1, II do 1, III do 5, IV do 2; tibiae: I & II spineless, III pl 3 rl 2 plv 3 rlv 2 vt 2, IV pl 3 rl 2 plv 2 rlv 2 vt 2; metatarsi: I spineless, II plv 1 rlv 1, III pl 3 rl 3 plv 2 rlv 3 vt 2, IV pl 2 do 3 rl 3 plv 4 rlv 2 vt 2; palpal spination: femora do 3; patellae rl 1; tibiae pl 4 do 2 rl 3 rlv 3; tarsi spineless. Abdomen dark-brown ( Fig. 98), orange at anterior margin, with six dense white setae markings; anterior markings closer together, middle markings widely separated, posterior markings moderately wide; lateral sigilla circular; venter light brown with orange background, with few brown feathery setae; white speckle above spinnerets on anal tubercle. Spinnerets pale. Epigyne with very shallow anterior hood, anterior margin slightly concave, posterior margin slightly convex, almost parallel-sides, lateral margins ending abruptly; lateral hoods slightly curved, near copulatory openings; ST II, ST I and connecting ducts orange; ST I smaller than ST II, ST II elongate and directly anterolaterally ( Figs 63, 123, 124).   Type material.  Holotype ♀:  SOUTH AFRICA:  Eastern Cape Province: Asante Sana Game Reserve, between Waterkloof and Zuurkloof, 32°16.920'S, 24°58.800'E,  13.X.2010, leg. J. Midgley( pitfall traps, grass tussocks) ( NCA 2012/4162).   Paratypes:  SOUTH AFRICA:  Eastern Cape Province: Asante Sana Game Reserve, between Waterkloof and Zuurkloof, 32°16.020'S, 25°00.240'E,  23.X.2010, leg. J. Midgley( pitfall traps, tussock grass), 1♀( NCA 2012/ 4161); Asante Sana Game Reserve, Waterkloof, 32°16.980'S, 24°56.340'E,  23.X.2010, leg. J. Midgley( pitfall traps), 1♀( NCA 2012/4160), 1♀( NCA 2012/4158).   Distribution.Known from the typelocality in the Eastern Cape, South Africa( Fig. 125). 2236308073 2010-10-13 NCA J. Midgley South Africa -32.282 Asante Sana Game Reserve 1 24.98 24 25 NCA 2012 1 1 Eastern Cape holotype 2236308066 2010-10-23 NCA J. Midgley South Africa Asante Sana Game Reserve -32.283 Waterkloof 1 24.939 Asante Sana Game Reserve 24 25 NCA 2012 3 3 Eastern Cape paratype