Frolov & Scholtz 2005 : 388 An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa Abdalla, Ishtiag H. Deschodt, Christian M. Scholtz, Clarke H. Sole, Catherine L. Zootaxa 2018 2018-10-23 4504 1 41 75 3X4TS Frolov & Scholtz, 2005 Frolov & Scholtz 2005 [151,756,1113,1139] Insecta Scarabaeidae Macroderes GBIF Animalia Coleoptera 22 63 Arthropoda species namakwanus  ( Figs. 13, 35, 42, 61, 75)     Macroderes namakwanus  Frolov & Scholtz 2005: 388.     Typelocality. RSA, Western Cape Prov, Hoekbaai, 31°09'26.07''S 017°45'55.70''E.  Type material examined.  Holotype, ♂( TMSA): “ RSA/ Western Cape Prov/ Hoekbaai/ A. Frolov & C. Deschodtleg [printed] // AF-0039 (19) /  4–11. IX. 2003/ S 31° 04' 26.07'' E 017° 45' 55.70''[printed] // HOLOTYPUS/  Macroderes namakwanus/ Frolov & Scholtz / 2003 [red label, printed]”;  Paratypes: 1♂( TMSA): “S. Afr/ Namaquland/ Brakrivermouth / 36°.06' S–17°.44' E [printed] //  25. 8. 1979// E– Y: 1597 / groundtraps / 63 days / Endrödy—Youngaleg [printed] // PARTYPUS /  Macroderes namakwanus/ Frolvo & Scholtz / 2003 [red label, printed]”;  1 ♂( TMSA): “S. AFR / Namaquland/ Hoekbaai/ 2 Km/ E NE / 31°. 11' S 17°. 47' E [printed] //  27. 8. 1979// E–Y: 1612 / white sand / night / Endrödy—Youngaleg [printed] // PARATYPUS/  Macroderes namakwanus/ Frolov & Scholtz / 2003 [red label, printed]”;  1 ♂( TMSA): “S. Afr/ Namaquland/ Kotzerus/ 30° 57' S–17°.50' E [printed] //  23. 8. 1979// E–Y: 1581 / white dune / day /. Endrödy—Youngaleg [printed] // PARATYPUS/  Macroderes namakwanus/ Frolov & Scholtz / 2003 [red– printed]”;  1 ♀( TMSA): “S. Afr/ Richtersveld/ Klein Helskloof/ 28°.51' S–17°.24' E [printed] //  8.9.1976// E–Y: 1239 / groundtraps / 32 days / Endrödy—Youngaleg [printed] // groundtrap with faeces bait [printed] // PARATYPUS/  Macroderes namakwanus/ Frolov & Scholtz / 2003 [red label, printed]”.  Additional material examined.  2 ♂♂and 3 ♀♀( UPSA): “ South Africa/ 6 kmN of / Kotzerus/ S 30° 69' 26.11'' E 017° 86' 9.83'' /  12. 8. 2015”.  Size range.Males length: 10.4–12.8 mm, width: 7.0– 8.7 mm; females length: 11.7–12.5 mm, width: 8.0–9.0 mm.   Differential diagnosis.  M. namakwanusis in habitus similar to  M. endroedyi( Fig. 13) however, it is differentiated by the combination of the following characteristics: Pronotum regularly punctate lacking the distinct tubercle in between ( Fig. 61) and no shiny tubercles on the elytra ( Fig. 75)   Habitat and distribution. See distribution under  M. mutilans( Fig. 2) 2028137807 [151,1081,1286,1311] South Africa -31.157242 Hoekbaai 1 17.765472 Prov 22 63 1 Western Cape holotype 2028137793 2003-09-04 2003-09-11 2003-09-04 TMSA C. Deschodt South Africa -31.07391 Hoekbaai 1 17.765472 Prov 22 63 1 1 Western Cape holotype 2028137809 1979-08-25 TMSA Endrody-Younga South Africa Afr Brakriver Namaquland 22 63 1 1 paratype 2028137797 1979-08-27 TMSA Endrody-Younga South Africa Namaquland Km Hoekbaai 22 63 1 1 paratype 2028137813 1979-08-23 TMSA Endrody-Younga South Africa Afr -30.95 Kotzerus Namaquland 22 63 1 1 paratype 2028137782 1976-09-08 TMSA Endrody-Younga South Africa Afr Klein Helskloof Richtersveld 22 63 1 1 paratype 2028137783 2015-08-12 UPSA South Africa Kotzerus 22 63 5 3 2