Penthetria Meigen (Diptera: Bibionidae): Revision of the New World species and world catalog Fitzgerald, Scott J. Zootaxa 2021 2021-02-10 4926 4 451 500 76Q42 Hardy, 1945 Hardy 1945 [151,549,1759,1786] Insecta Bibionidae Penthetria Animalia Diptera 16 467 Arthropoda species distincta  ( Figs. 9–11, 42, 48, Map 1)   Type Material.   Holotypemale(not examined):  GUATEMALA: “Antigua,”  24 June 1923, E.G. Smyth( USNM).   Paratypes(examined):  GUATEMALA: GUATEMALA: Guatemala City,  2 May 1923, E.G. Smyth, 4M ( USNM),   20 July 1923, 1M ( USNM);  SACATEPÉQUEZ: “ Antiqua,” D.G. Eisen, 2M ( USNM); “ Antiqua,”  24 June 1923, E.G. Smyth, 1F ( allotype) ( USNM);  QUETZALTENANGO: Volcan Santa María, Nov., Schaus& Barnes, 1M ( USNM).   FIGURES 9–11.  P. distincta, male terminalia. 9. Dorsal. 10. Posterior. 11. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine; vmc, ventromedian cleft of gonocoxite; vml, ventromedian lobes of gonocoxite.  Additional Material examined.   EL SALVADOR: LA LIBERTAD: Santa Tecla, 7 14 95, P. A.B., 1M, 1F ( USNM);   GUATEMALA: GUATEMALA: Puerta Parada,  1850 m, Bosquescipres, J.C. Schuster,  15–22 XII 2012, 1M ( UVGC),   22–29 XII 2012, 2M ( SFC),   29 XII 2012–  5 I 2013, 1F ( SFC),   22 II–1 III 2014, 1F ( SFC);  Puerta Parada,  1850 m, Casa Jack, 14.556630, -90.463362,  5–12 Jan 2013, J.C. Schuster, 1F ( SFC);  Puerta Parada,  1840 m,  7–14 VII 2018, J.C. Schuster, 1F ( UVGC),   14–21 VII 2018, 1M ( SFC);  TOTONICAPÁN: Momostenango,  9 Dec. 1979, 1M ( UCDC);   MEXICO: CHIAPAS:  7200 ft., S. Crist.las Casas,  1 June 1969, Malaise trap, 5M, 1F ( CNCI);  Sn. Cristobal,  22.VII.1969, L.A. Kelton, 1M ( CNCI).   Diagnosis. Males of  P. distinctacan be distinguished from all New World congeners by the following combination of characters: tergite nine with posteromedial emargination deep (nearly subdividing the tergite) ( Fig. 9) with the apices of the resulting lobes truncate and shelf-like (best observed in posterior view) ( Fig. 10), posteroventral edge of tergite nine with a field of densely-set, short, black, spine-like setae, which contrast the typical setae on the remainder of the tergite (difficult to see in undissected specimens), gonostylus shining, basally with long hairs, but more distally with only very minute and spare hairs, slender and apically coming to a narrowly rounded point ( Figs. 9–11), ventromedial lobes of gonocoxite absent to very weakly developed ( Fig. 11), and R 2+3often with a basal appendix (as in Fig. 2).   Remarks.  P. distinctais a fairly large species with male wing 7.0–9.5 mm (n=4) and female wing 12.0–12.5 mm (n=2) and is most similar to  P. appendicula(see “Remarks” under  P. appendicula).  Geographic & Seasonal Distribution. Guatemalaand Mexico( Map 1); additional Guatemalan records in Hardy (1945). Seasonal distribution summarized in Table 1. 3037447436 [335,1433,1868,1895] 1923-06-24 USNM E. G. Smyth Guatemala GUATEMALA 16 467 1 holotype 3037447359 [151,1432,1904,1931] 1923-05-02 USNM E. G. Smyth Guatemala Guatemala City 16 467 1 Guatemala paratype 3037447461 [151,466,1940,1966] 1923-07-20 USNM Guatemala 16 467 1 paratype 3037447518 [479,1132,1940,1968] USNM D. G. Eisen Guatemala Antiqua 16 467 1 Sacatepequez paratype 3037447306 1923-06-24 USNM E. G. Smyth Guatemala Antiqua 16 467 1 allotype 3037447391 USNM Schaus & Barnes Guatemala Volcan Santa Maria 16 467 1 Quetzaltenango paratype 3037447440 USNM A. Brune El Salvador LA LIBERTAD 18 469 1 LA LIBERTAD 3037447397 2012-12-15 2012-12-22 2012-12-15 UVGC J.C. Schuster Guatemala 1850 GUATEMALA 18 469 1 Guatemala 3037447304 [313,632,223,249] 2012-12-22 2012-12-29 2012-12-22 SFC Guatemala 18 469 1 Guatemala 3037447564 [646,1024,222,249] 2012-12-29 2013-01-05 2012-12-29 SFC Guatemala 18 469 1 Guatemala 3037447547 [1038,1351,223,249] 2014-02-22 2014-03-01 2014-02-22 SFC Guatemala 18 469 1 Guatemala 3037447368 2013-01-05 2013-01-12 2013-01-05 SFC J. C. Schuster Guatemala 1850 14.55663 1 -90.46336 18 469 1 Guatemala 3037447467 2018-07-07 2018-07-14 2018-07-07 UVGC Puerta Parada & J. C. Schuster Guatemala 1840 18 469 1 Guatemala 3037447326 [639,950,295,321] 2018-07-14 2018-07-21 2018-07-14 SFC Guatemala 18 469 1 Guatemala 3037447457 1979-12-09 UCDC Momostenango Guatemala TOTONICAPAN 18 469 1 Totonicapan 3037447556 [377,1432,330,357] 1969-06-01 CNCI Crist. & Casas Mexico 2195 CHIAPAS 18 469 1 Chiapas 3037447463 [151,746,367,393] 1969-07-22 CNCI Sn. Cristobal & L. A. Kelton Mexico 18 469 1 Chiapas