A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini) Silva, Fernando A. B. Valois, Marcely Zootaxa 2019 2019-07-08 4629 3 301 341 4W3LQ Silva & Valois Silva & Valois 2019 [151,651,898,925] Insecta Scarabaeidae Scybalocanthon Animalia Coleoptera 25 326 Arthropoda species martinezi sp. nov.  ( Figs. 1P, 4O, 5N, 6D)   Etymology.Named in honour of Antonio Martínez, for his description of  Scybalocanthonand his valuable contributions towards the taxonomy of the genus.   Diagnosis. Specimens of  S. martineziare similar to those of  S. arnaudi,  S. federicoescobari,  S. papaxibe,  S. pinopterus, and  S. uniplagiatusby having the pronotum uniformly colored, or with one elliptical spot on the central portion ( Figs. 1P, 2A, 1I, 2K, 1D, 1T); femora almost completely yellow or brown, with black spots only on the tips ( Fig. 3G); eighth elytral stria with thin carina at the anterior portion ( Fig. 3D); endophallus with bristles right beside the FLP sclerite ( Figs. 5N, R, H, W, D, Q); and additional sclerite ( AS) absent ( Figs. 5N, R, H, W, D, Q). They can be distinguished from those of  S. pinopterusand  S. arnaudiby the strongly asymmetrical parameres; left paramere with acute projection on dorsal margin and bilobate excavation on ventral margin ( Fig. 4O) (other species with slightly asymmetrical parameres, lacking acute projection and bilobate excavation on dorsal and ventral margins, respectively ( Figs. 4S, E)); from those of  S. federicoescobari,  S. papaxibe, and  S. uniplagiatusby the bilobate excavation of ventral margin of left paramere wide and deep, extending at least one-half of the paramere margin in the lateral view ( Fig. 4O) (other species with a short bilobate excavation, not reaching one-half of paramere margin in the lateral view ( Figs. 4I, W, R)).   Description. Body.Oval-elongate. Surface opaque (except pronotum), completely microgranulate. Color. Most of pronotum, hypomera (except internal margin), metaventrite, abdominal ventrites, pygidium, middle of femora, and tibiae yellow or light brown. Head, internal margin of hypomera, prosternum, mesoventrite, mesoepisternae, metaepisternae, elytra, trochanters, and proximal and distal portions of femora dark brown. Length. 8.5 mm. Thorax. Pronotum with one brown rounded spot at the anterocentral portion ( Fig. 1P). In some, spot absent. Anterior angles of pronotum approximately 80°. Lateral margin regularly curved outward, not forming an angle at the middle portion. Elytra. Striae thin and shiny, punctures conspicuous. Eighth stria with a thin carina at the anterior portion. Aedeagus.Parameres strongly asymmetrical ( Fig. 4O). Dorsal margin of right paramere curved inward, apex obliquely truncate. Ventral margin of right paramere substraight, with a rounded excavation at the basal portion. Dorsal margin of left paramere curved inward from the basal to medial portions, apical portion with a short and pointed projection. Ventral margin of left paramere substraight, with a bilobate excavation from the basal to medial portions. SRP circular, with rounded handle-shaped extension ( Fig. 5N). FLP short, comma shaped, with three sets of bristles right beside it ( Fig. 5N). A+SA with two superposed and elongate sclerites ( Fig. 5N).   Type material.   Holotype. ECUADOR: NAPO: Daimi1, Bosque Sendero,  20.ix.1989, pitfall, Sandoval— 1♂( CEMT).   Paratypes[ 19 males, 18 females].  ECUADOR: NAPO: Scyasuni,  20–21.x. 1997, 200 m, D. Padilla& I. Tapin— 1♀( CEMT).   COLOMBIA: AMAZONAS, Leticia,  215 m( 4°11’20’’S, 69°56’9’’W),  10.vii.1970, H. & A. Howden— 3♂ 1♀( CMNC); Leticia,  215 m( 4°11’20’’S, 69°56’9’’W),  19–25.ii.1972, H. & A. Howden— 1♀( CMNC); Leticia,  215 m( 4°11’20’’S, 69°56’9’’W),  20–25.ii.1972, Peck& Howden, dung traps— 10♂ 4♀( CMNC, MZUFPA); Los Alpes, Leticia,  213 m( 4°4’25’’S, 70°0’5’’W), 23.ii–iii.1974, S. Peck, dung trap— 2♂ 10♀( CMNC); road end, Leticia,  213 m( 4°4’9’’S, 69°59’56’’W),  23.ii–2.iii.1974, S. Peck, dung trap— 4♂ 1♀( CMNC).   Distribution.Known from Ecuadorand Colombia( Fig. 6D). Endemism areas: Brazilian sub-region: Boreal Brazilian dominion: Napo and Imerí provinces (see Morrone 2014; fig. 12). 2847261398 1989-09-20 CEMT Ecuador Bosque Sendero Daimi 25 326 1 1 Napo holotype 2847261568 1997-10-20 1997-10-21 1997-10-20 CEMT D. Padilla Ecuador 26 327 Scyasuni 25 326 38 19 19 Napo paratype 2847261312 1970-07-10 1974-03-02 1970-07-10 CMNC, MZUFPA Peck & Howden & S. Peck Colombia 215 -4.0691667 Leticia 21 -69.998886 26 327 36 17 19 paratype 2847261504 1970-07-10 1974-03-02 1970-07-10 CMNC, MZUFPA Peck & Howden & S. Peck Colombia 215 -4.0691667 Leticia 21 -69.998886 26 327 36 17 19 paratype 2847261346 1970-07-10 1974-03-02 1970-07-10 CMNC, MZUFPA Peck & Howden & S. Peck Colombia 215 -4.0691667 Leticia 21 -69.998886 26 327 36 17 19 paratype 2847261410 [284,1436,260,287] 1970-07-10 1974-03-02 1970-07-10 CMNC, MZUFPA Peck & Howden & S. Peck Colombia 215 -4.0691667 Leticia 21 -69.998886 26 327 36 17 19 paratype 2847261488 [151,1328,296,323] 1970-07-10 1974-03-02 1970-07-10 CMNC, MZUFPA Peck & Howden & S. Peck Colombia 215 -4.0691667 Leticia 21 -69.998886 26 327 36 17 19 paratype