Studies in the Liocranidae (Araneae): revision of Andromma Simon, 1893 Bosselaers, Jan Jocqué, Rudy European Journal of Taxonomy 2022 2022-12-09 850 1 78 Bosselaers & Jocqué, 2022 Bosselaers & Jocqué 2022 [603,887,264,291] Arachnida Liocranidae Andromma Animalia Araneae 46 47 Arthropoda species ghesquierei sp. nov.   Figs 2E, 24–25, 40    Diagnosis The male of  Andromma ghesquierei sp. nov.differs from the males of all other known species of  Andrommaby the male palp with the embolus originating in the middle of the posterior tegular margin, running over the centre of the tegulum ( Fig. 24C–D); the female can easily be recognized by the very large, rounded spermathecae visible in transparency ( Fig. 25C–D).    Etymology The species name is a patronym in honour of J. Ghesquière who collected all the typematerial of this species apart from the specimens of Lac Tumba; he also collected many other invertebrates in D.R. Congo.    Type material   Holotype  D.R. CONGO • ♂; Equateur Province, Eala; 0°30′ N, 18°19′ E; 1936; J. Ghesquièreleg.; BE_ RMCA_ ARA.Ara 247536.   Paratypes  D.R.  CONGO• 18 ♂♂, 14 ♀♀, 3 juvs; same collection data as for holotype; BE_ RMCA_ARA.Ara 177019•  1 ♀;  Nov. 1936; further same collection data as for holotype; BE_ RMCA_ARA.Ara 894•  1 ♀; same collection data as for preceding; BE_ RMCA_ARA.Ara 916•  3 ♀♀, 2 juvs;  May 1936; further same collection data as for preceding; BE_ RMCA_ARA.Ara 22957•  1 ♀; same collection data as for preceding; BE_ RMCA_ARA.Ara 917•  1 ♀; same collection data as for preceding; BE_ RMCA_ARA.Ara 920•  1 ♀; same collection data as for preceding; BE_ RMCA_ARA.Ara 921•  1 ♀;  EquateurProvince, Lac Tumba, Bikoro; 0°43′ S, 18°07′ E;  350 ma.s.l.;  11 Oct. 1955; N. Leleupleg.; BE_ RMCA_ARA.Ara 84109.   Other material  D.R. CONGO• 16 ♂♂, 7 ♀♀; N Kivu., Rutshuru; 1°11′ S, 29°27′ E;  20 Dec. 1937; J. Ghesquièreleg.; BE_RMCA_ARA.Ara 22878.    Description  Male( holotype, Fig. 24) MEASUREMENTS. Total length 3.20. Carapace length 1.42, width 1.21, height 0.64. COLOUR ( Fig. 24A–B). Cephalothorax including legs and chelicerae yellowish orange, fovea brownish; sternum pale yellow with small brown precoxal triangles; abdomen including spinnerets pale cream. PROSOMA. Fovea faint, length 0.21, anterior end 0.71 from PME. MOQ ( Fig. 2E) length 0.18, anterior width 0.20, posterior width 0.19. AER straight from above, width 0.32, PER procurved from above, width 0.35. AME circular, other eyes oval; AME 0.08, ALE 0.05, PLE 0.05, PME 0.02; AME–AME: 0.03, AME–ALE: 0.01, ALE–PLE: touching, PME–PME: 0.12. Clypeus vertical, 0.06, with four long setae. Chilum absent. Sternum 0.92 long, 0.78 wide. Chelicerae: anterior margin with three teeth. LEG SPINATION. All femora with one dorsal spine, Fe I pl 1; T I v 2-2-2-2-2; Mt I v 2-2-2; T II v 2-2-2-2; Mt II v 2-2-2; T III v 2-2; Mt III pl1v 2; T IV pl 1 v 1; Mt IV rl 1 v 1-2.   Fig. 24.  Andromma ghesquierei sp. nov., ♂, holotype (BE_RMCA_ARA.Ara 247536). A. Habitus, dorsal view. B. Same, ventral view. C–D. ♂, left palp, ventral view. E–F. Same, retrolateral view.Abbreviations: CD= copulatory duct; CO=copulatory opening; FD =fertilisation duct; ST =spermatheca. Scale bars: A–B =1 mm; C–F =0.2 mm. LEG I MEASUREMENTS. Fe: 1.07, P: 0.43, T: 0.85, Mt: 0.78, t: 0.64; total: 3.77. LEG FORMULA. 4123. MALE PALP ( Fig. 24C–F). RTA subcircular, provided with thin prong pointing forward, VTA subtriangular with rounded corners, both in retrolateral and ventral views; tegulum with sperm duct long and complex with wide basal part clearly visible in ventral view; embolus with thick base, originating in centre of posterior margin of tegulum, curved inward, ending in slender tip slightly curved antero-ventrally, near anterior margin of cymbial concavity. Conductor close to distal tip of embolus, subrectangular with a prolateral, forward projecting finger-like appendage.  Female( ParatypeBE_RMCA_ARA.Ara 22957; Fig. 25) MEASUREMENTS. Total length 3.69. Carapace length 1.56, width 0.99, height 0.71. COLOUR ( Fig. 25A–B). Carapace and chelicerae pale yellowish orange, legs pale yellow, sternum yellow with pale brown precoxal triangles. PROSOMA. Fovea pronounced, length 0.28, anterior end 0.71 from PME. MOQ length 0.25, anterior width 0.25, posterior width 0.26. AER straight from above, width 0.40, PER procurved from above, width 0.45. AME circular, remainder oval; AME 0.10, ALE and PLE 0.05, PME 0.02; AME–AME: 0.05, AME–ALE: 0.03, ALE–PLE: touching, PME–PME: 0.16. Clypeus 0.07. Chelicerae with three teeth. Chilum absent. Sternum 0.92 long, 0.78 wide. LEGS: TARSI SLIGHTLY WIDENED TOWARDS EXTREMITY. Spination: all femora with one dorsal spine; T I v 2-2- 2-2-2; Mt I v 2-2-2; T II v 2-2-2-2; Mt II v 2-2-2; T III v 2-2; Mt III v 2-2; T IV pl 1 v 1; Mt IV rl 1 v 1-2.  LEG MEASUREMENTS.     Fe P T Mt t total  I 1.47 0.49 1.33 1.33 0.98 5.60  II 1.40 0.49 1.05 0.77 0.70 4.41  III 1.12 0.49 0.91 0.98 0.84 4.34  IV 1.47 0.56 1.26 1.47 0.98 5.74 LEG FORMULA. 4123. EPIGYNE ( Fig. 25C–F). Roughly rectangular, slightly wider than long, with crescent-shaped copulatory openings in the posterior half. Large broadly oval spermathecae and copulatory ducts visible in transparency. Fertilisation duct short, oriented posteriorly. Sickle-shaped sclerites procurved.    Distribution Known from the Eastern D.R. Congoand the Kivu Province( Fig. 40).    Remark It would be surprising that the species occurs in the eastern lowland of Congoas well as in the western Kivu Province. The specimens from Kivumay be mislabeled. 3990057310 1936 BE_RMCA_ ARA J. Ghesquiere 0.5 Eala 1308 18.316668 46 47 247536 1 1 CONGO holotype 3990057325 BE_RMCA_ARA J. Ghesquiere & Ara 0.5 CONGO 1308 18.316668 46 47 177019 32 14 18 CONGO holotype 3990057352 [307,1378,1057,1084] 1936-11 BE_RMCA_ARA J. Ghesquiere & Ara 0.5 Ara 1308 18.316668 46 47 894 1 1 Bas-Congo holotype 3990057309 [189,1052,1094,1121] 1936-11 BE_RMCA_ARA J. Ghesquiere & Ara 0.5 Ara 1308 18.316668 46 47 916 1 1 Bas-Congo paratype 3990057346 1936-05 BE_RMCA_ARA J. Ghesquiere & Ara 0.5 Ara 1308 18.316668 46 47 22957 3 3 Bas-Congo paratype 3990057349 1936-05 BE_RMCA_ARA J. Ghesquiere & Ara 0.5 Ara 1308 18.316668 46 47 917 1 1 Bas-Congo paratype 3990057319 1936-05 BE_RMCA_ARA J. Ghesquiere & Ara 0.5 Ara 1308 18.316668 46 47 920 1 1 Bas-Congo paratype 3990057338 [536,1379,1204,1230] 1936-05 BE_RMCA_ARA J. Ghesquiere & Ara 0.5 Ara 1308 18.316668 46 47 921 1 1 Bas-Congo paratype 3990057313 1955-10-11 BE_RMCA_ARA N. Leleup Lac Tumba 350 -0.71666664 Bikoro 1308 18.116667 Bikoro 46 47 84109 1 1 Equateur Province paratype 3990057329 1937-12-20 BE_RMCA_ARA J. Ghesquiere Republic of the Congo -1.1833334 Rutshuru 1308 29.45 46 47 22878 23 7 16 Kivu