Encephalus angusticollis J. Sahlberg 1880: 82
Encephalus solskyi Heyden 1880: 68
Solsky 1875 : 270
G. nitidula (Gyllenhal 1810))
Eppelsheim 1893 : 39
Pace 1985 : 94
Pace 1989 : 486
Review of the subtribe Gyrophaenina Kraatz 1856 (Coleoptera: Staphylinidae: Aleocharinae: Homalotini) of Middle Asia, with emphasis on the fauna of Kazakhstan
Enushchenko, Ilya V.
Zootaxa
2019
2019-06-05
4613
2
379
397
39NFS
J. Sahlberg 1880
J. Sahlberg
1880
[151,841,1413,1440]
Insecta
Staphylinidae
Encephalus
Animalia
Coleoptera
1
380
Arthropoda
species
angusticollis
Encephalus
( Figs. 1–12)
Encephalus angusticollisJ. Sahlberg 1880: 82 Encephalus solskyiHeyden 1880: 68(replacement name of Encephalus kraatzi Solsky 1875: 270; not synonym of G. nitidula(Gyllenhal 1810)) resyn. Encephalus torosus Eppelsheim 1893: 39 syn. nov. Encephalus torosus: Pace 1985: 94 Encephalus(s.str.) torosus: Pace 1989: 486
Type material examined: Holotypeof Encephalus angusticollisJ. Sahlberg 1880, ♂: ‘Jeniseisk’, ‘J. Sahlb.’, ‘J. Sahlb. | ind. typ.’ ˂all labels rectangular, printed˃, ‘1135.’ ˂yellow rectangular label, handwritten˃, ‘20’ ˂square label, printed˃, ‘…’ ˂empty square red label˃, ‘ Encephalus| angusticollis| J. Sahlb.’ ˂rectangular label with black frame, handwritten˃, ‘ZMUT | TYPE538’ ˂rectangular red printed label˃ (ZMUT) ( Figs. 4–6). Labels as in Fig. 6. Lectotypeof Encephalus torosus Eppelsheim 1893, ♂[a plastic plate with a preparation of the aedeagus, VIII male tergite and sternite in Canadabalsam]: ‘c. Eppelsh. | Steind. d.’ [rectangular, printed label], ‘ torosus| Epp.’ [rectangular, handwritten label], ‘Ost-Sibirien. |Auchlyebiet | Ecs Irkut. | Leder 1891’ [handwritten label], ‘ ♂’ [little, square, handwritten label], ‘ torosus| Epp. Deutschient. | Zeit. 1983. p. 39’ [handwritten label], ‘TYPUS’ [red rec- tangular label, printed], ‘ Encephalus| torosusEpp.| det. R. Pace 1984 | Lectotypus’ [white rectangular label; mostly handwritten] (NHMW) ( Figs. 7–9). Labels as in Fig. 9. Paralectotypeof Encephalus torosus Eppelsheim 1893, 2 ♀♀: [a plastic plate with a preparation of the sper- matheca in Canadabalsam]: ‘c. Eppelsh. | Steind. d.’ [rectangular, printed label], ‘ torosus| Epp.’ [rectangular, handwritten label], ‘Ost-Sibirien. | Auchlyebiet | Ecs Irkut. | Leder 1891’ [handwritten label], ‘TYPUS’ [red rectan- gular label, printed], ‘ Encephalus| torosusEpp.| det. R. Pace 1984 |’ [white rectangular label; mostly handwritten] (NHMW) ( Figs. 10–12). Labels as in Fig. 12. Materialexamined: KAZAKHSTAN: East Kazakhstan Area: Katon-Karagaiskiy District: 3 ♂♂, 21 ♀♀: Altai Mountains, Ustʼ-Kamenogorsk [=Oskemen], Rakhmaninovskie Klyuchi, Berelʼ River, 21.VII.1989, V.A. Kashcheevleg. ( ZIN, cIE) ( Figs. 1–3). Additional material examined: RUSSIA: Khakasia: Altaiskiy District: ♂: Abakan, left side of YeniseiRiv- er, in fungi on the willow trunk, 3.VII.1988, E.M. Veselovaleg. ( ZMM); Evenk Autonomous Region: Balkitskiy District: ♂, 2 ♀♀: Podkamennaya Tunguska Riverbasin, Kochumbek River, 8 kmlower the mouth, № 267, 24.VIII.1990, A.B. Ryvkinleg. (cIE); Krasnoyarsk Territory: Turukhanskiy District: ♀: Nizhnyaya Sarchikha River, litter under bracket fungus on birch, 11.VII.1992, V.B. Semenovleg. ( ZMM); ♀: Bol’shaya Varlamovka River, 6 kmlower Malaya Rivermouth, 25.VIII.1988, A.B. Ryvkinleg. ( ZMM); Tyva: Tandinskiy District: 2 ♀♀: Durgen, 4.VII.1988, E.M. Veselovaleg. ( ZMM); Irkutsk Area: Slyudyanskiy District: ♀: Baikalʼsk, Solzan valley, 24.VI.1978, V.G. Shilenkovleg. (cIE); Buryatiya: Tunkinskiy District: 3 ♂♂, 2 ♀♀: Arshan, right side of Kyngyrga River, 26–29.VIII.2009, I.V. Enushchenkoleg. (cIE); Zabaykalsky Krai: Uletovskiy District: ♂: Lukovaya River(right tributary of Ingoda River) valley, N 49°41’83,8’’, E 111°08’59,5’’, h= 1420 m, 22.VII.2009, A.V. Shavrin& I.V. Enushchenkoleg. (cIE); Krasnochikoiskiy District: ♀: valley of Kunalei River, 6 kmNW Kunalei, 21.VIII.1998, A.V. Shavrinleg. (cIE); Kyrinskiy District: ♂, 2 ♀♀: Bukukun River, N 49°27’17,6’’, E 111°08’15,6’’, h= 1137 m, 6.VIII.2013, I.V. Enushchenkoleg. (cIE); Khabarovsk Territory: Verkhnebureinskiy District: ♀: Dublikanskiy Nature Refuge, right side of Dublikan River valley, 1 kmN cordon, mosses and plant debris on open sedge hummocky with grasses and bushes, 2.IX.2009, A.B. Ryvkinleg. ( ZMM); Amur Area: Skovorodinskiy District: ♂, 2 ♀♀: left side of Urka River, ca. 4 kmN Yerofei Pavlovich, N 54°00’40,89’’, E 121°57’43,75’’, h= 525 m, 1.IX.2014, A.V. Shavrin& I.V. Enushchenkoleg. (cIE); Mazanovskiy District: ♀:, Nora Riverbasin, Sorokavyorstnaya chan- nel, foot of Sosnovaya mountain, 215 ma.s.l., 27.07.2006. E.M.Veselova& A.B.Ryvkinleg. (cIE); MONGOLIA: Tuve[ Tzentralnyi] Aimak: ♂, ♀: Zaisan, Sslope of Bogdo-Ula Mountain, 4.VII.1967, I.M. Kerzhnerleg. ( ZIN).
Remarks. J.Sahlberg (1880) described E. angusticollisbased on a single male from ‘... territorio silvosoprope oppidum Jenisseisk…’ (now known as Yeniseysk in Krasnoyarsk Territory, West Siberia). Eppelsheim (1893)de - scribed E. torosus( Figs. 7–12) from south-western Cisbaikalia, based on ‘3 oder 4 Stücke’. The author compared this species with E. angusticollisand E. complicans, from which he distinguished them by the distinctly broader body and coarser microsculpture. Pace recorded E. torosusfrom Nepal( Pace 1985, 1989) and revised types ( 2 males, 2 females) of E. torosusfrom NHMW ( Pace 2006), without designation of a lectotype. I have revised these types and found that, based on the morphological details of the body, these specimens are conspecific with E. angusticollis. It is worth noting that the possible synonymy of E. torosuswith E. angusticolliswas assumed by Pace (1985). Thus, a new synonymy is proposed here. After the descriptions of Encephalus kraatziby Hochhuth (1872)from Ukraineand Solsky (1875)from the Baikal Region, there was a very complicated story with these identical names discussed in Enushchenko & Shavrin (2011). Encephalus solskyiL.Heyden 1880(replacement name of E. kraatzi Solsky 1875) was synonymized with G. nitidula(Gyllenhal 1810), while E. kraatzi Hochhuth 1872was transferred to the genus Brachidaby Shilov (1977)and synonymized with B. exiguaHeer 1839. According to the original description of E. kraatziby Solsky (1875), the species is characterized by transverse antennomeres as in specimens of Encephalus(‘Antennis crassiusculis, articulis penultimis transcersis…’), whereas European G. nitidulacan be distinguished by elongate antennomeres. Thus, I propose that E. solskyiHeyden 1880is not the same as G. nitidulaand should be resynonymized with E. angusticollis( Figs. 4–6). This species is recorded from Kazakhstanand Middle Asia for the first time.
2248084560
1989-07-21
ZIN
V. A. Kashcheev
Kazakhstan
Area
Rakhmaninovskie Klyuchi
Altai Mountains
2
381
24
21
3
Shyghys Qazaqstan Oblysy
2248084598
1988-07-03
1990-08-24
1988-07-03
ZMM
Abakan & Yenisei & E. M. Veselova & A. B. Ryvkin
Russia
Altaiskiy District
Balkitskiy District
Evenk Autonomous Region
2
381
4
2
2
Khakasiya
2248084557
1988-08-25
1992-07-11
1988-08-25
ZMM
Territory & V. B. Semenov & A. B. Ryvkin
Russia
Turukhanskiy District
Bol'shaya Varlamovka River
Nizhnyaya Sarchikha River
2
381
2
2
Krasnoyarsk
2248084586
1988-07-04
ZMM
E. M. Veselova
Russia
Durgen
Tandinskiy District
2
381
2
2
Tyva
2248084571
1978-06-24
V. G. Shilenkov
Russia
Area
Baikal'sk
Slyudyanskiy District
2
381
1
1
Irkutsk
2397873914
2009-08-26
Russia
2
381
Buryatiya
2397873911
2009-07-22
Russia
49.683334
111.14986
2
381
Buryatiya
2397873913
1998-08-21
Russia
2
381
Buryatiya
2397873912
2013-08-06
Russia
49.45489
111.137665
2
381
Buryatiya
2248084569
2009-09-02
ZMM
Dublikanskiy Nature Refuge & A. B. Ryvkin
Russia
Territory
Dublikan River valley
Verkhnebureinskiy District
2
381
1
1
Khabarovsk
2248084563
2006-07-27
2014-09-01
2006-07-27
Area & A. V. Shavrin & I. V. Enushchenko & E. M. Veselova & A. B. Ryvkin
Russia
Skovorodinskiy District
525
54.01136
Mazanovskiy District
1
121.96215
Urka River
2
381
4
3
1
Amur
2248084597
1967-07-04
ZIN
Tzentralnyi & Zaisan, S & I. M. Kerzhner
Mongolia
Bogdo-Ula Mountain
Aimak
2
381
2
1
1
Tov