Bolívar, 1890 : 232 Karsch 1893 : 204 Kirby 1906 : 106 Giglio-Tos 1907 : 26 Bolívar 1910 : 544 Chopard 1954 : 80 Chopard 1968 : 349 Robillard & Desutter-Grandcolas 2004a : 578 Robillard et al. 2007 : 1267 Nattier et al. 2011 : 2201 Chintauan-Marquier et al. 2016 : 62 Vicente et al. 2017 : 2203 Orthoptera Xenogryllus Kevan & Knipper 1961 : 368 Chopard 1968 : 349 Taxonomic revision of the genus Xenogryllus Bolívar, 1890 (Orthoptera, Gryllidae, Eneopterinae, Xenogryllini) Jaiswara, Ranjana Dong, Jiajia Ma, Libin Yin, Haisheng Robillard, Tony Zootaxa 2019 2019-01-18 4545 3 301 338 7G8LM Bolivar, 1890 Bolivar 1890 [151,642,1244,1270] Insecta Gryllidae Xenogryllus GBIF Animalia Orthoptera 5 306 Arthropoda species eneopteroides  ( Figs 1A–C, 3A–B, 4A, 5A, 6, 7A–C, 8A–B, 9A, 10A, 11A–B)     Xenogryllus eneopteroides  Bolívar, 1890: 232—  Karsch 1893: 204;  Kirby 1906: 106;  Giglio-Tos 1907: 26;  Bolívar 1910: 544;  Chopard 1954: 80;  Chopard 1968: 349; Chiffaud & Gillon 1984 (life history); Paris 1994[1993]: 69 ( lectotypedesignated);  Robillard & Desutter-Grandcolas 2004a: 578; 2004b: 273 (morphological phylogeny); 2006: 644; 2008: 67; 2011: 637; Robillard 2006: 673;  Robillard et al.2007: 1267(song frequency);  Nattier et al.2011: 2201(molecular phylogeny);  Chintauan-Marquier et al.2016: 62(molecular phylogeny);  Vicente et al.2017: 2203(historical biogeography); Cigliano et al.2018( OrthopteraSpecies File Online).    Xenogrylluseucopteroides—  Kevan & Knipper 1961: 368(misspelling);  Chopard 1968: 349.   Type material.   Lectotype, ♂,  Angola: Duque de Bragança, Bayão(Paris 1994) [ MNCN, examined on photograph].  Additional material examined.  Benin: Bas-Dahomey, plateaux de Zagnamadoet de Ketou, forêt d’Aquaqueré(saison sèche), 1910 P. Ducorps, 1♀(MNHN).   Cameroun:[L.] Conradt, 1♂, identified  Xenogryllus eneopteroidesBol.( MNHN).  Reg[ion] de Kribi, 1925, littoral, saison humide, Dr. Gromier, 1♂( MNHN).  Kamerun[Cameroun], Pipindi[Bipindi], Eitelded. 1♀, identified  Xenogryllus eneopteroidesBol.( MNHN).  10 kmN. of Yaoundé, E. of Agronomy Station, Nkolbisson,  10.vii.1975, N. D. Jago, 1♂, identified  Xenogryllus eneopteroidesBolivarby B. C. Townsend, 1976 ( NHMUK 010926525).   Central African Republic: République Centrafricaine,  75 kmWNW Mbaiki, N’Gotto[Ngoto], W Lobaye, savane arbustive,  7.iv.1995, nuit [night], L. Desutter-Grandcolas, 1 juvenile, n°14, molecular sample Xen-CA (MNHN-E0-ENSIF3159); 1♂, #1, sur plante [on plant] ( MNHN).   Democratic Republic of the Congo: Congo, 20 kmfrom L. Tumba(towards Coquilhatville) [Mbandaka],  5.iii.1964, from grassy clearing in forest, 1♂, 1♀, Tyson Roberts, Brit. Mus.1982-71 ( NHMUK 0 10926531, 010926572). Kalembe, 15.ix.1947, Miss. Tanganika, 1♀(RBINS).  Lukafa[Lukafu], Congo Belge,  6.xii.1938, H.J. Brédo, 1♀, R.Mus.Nat. Belg.I.G.12.204 ( RBINS).  Belgian Congo, Aru,  iii.1936, 5♂, 2♀, H.J. Bredo, dry Acaciabush ( NHMUK0 10926536, 0 10926527, 0 10926578, 0 10926580, 0 10926539, 0 10926566, 010926569). Niangara, v-vi-1915, 28°0 E 3°40’N, 1♂( AMNH).  Haut-Ituri: Faradje, 1♂, Blommaert, Musée du Congo, identified  Xenogryllus eneopteroidesby L. Chopard, R. Det.N 2814 ( MRAC).  S.E. Katanga: Ngaye: xixii.1931, 1♂, R.P. Claquin, Musée du Congo, identified  Xenogryllus eneopteroidesby L. Chopard, R. Det.N 2814 ( MRAC); 1932, 1♂, 1♀, R.P. Claquin, Musée du Congo, identified  Xenogryllus eneopteroidesby L. Chopard, R. Det.DD3509 ( MRAC).  Haut-Uele,  vi.1925, 1♀, Musée du Congo, identified  Xenogryllus eneopteroidesby L. Chopard, R. Det.N 2814 ( MRAC).  Kibali-Ituri: Geti, ii–iv.1937, 1♂, Ch. Scops, Musée du Congo, identified Xenogryllus eneopteroidesby L. Chopard, R. Det.B 5318 ( MRAC).   Gabon: Plateaux Batéké, piste Ekalla,  30 kmW Leconi[Lekoni], milieu herbeux,  19.vi.1994, L. Desutter-Grandcolas: 1 juvenile, n°61, jour [day], litière [leaf litter], molecular sample Xen-GA (MNHN-EO-ENSIF3442); 1♂, n°3, nuit [night], sur plante—base tronc [on plant, low on tree trunk] ( MNHN).  Bas Ogooué, 1♂, identified  Xenogryllus eneopteroidesBol.by L. Chopard( MNHN).  Ntoum,  19.vii.[19]85, A. Pauly, 2♂, 2♀, 2 juveniles, lumière [at light] ( RBINS).   Ghana: Eastern region, Hansowodze, 2 mls from Kade A.R.S.[Agricultural Research Station],  19.vii.1963, I. K. Bacheampong, 4♂( NHMUK0 10926524, 0 10926567, 0 10926528, 010926571). Ghana, Togoland plateau, grassy, summit above [Nkonya] Wurupong,  23.xii.1959, N.D. Jago, 1♂, Brit. Mus.1965-474 ( NHMUK 010926535).  Eastern region, Kade Agr. Res. Stn[Agricultural Research Station],  11.vii.1963, I. K. Bacheampong, 2♂, Brit. Mus.1965-474 ( NHMUK0 10926526, 010926538). Transvolta, Togoland, Amedzofe:  3.v.1959, montane grass, 1♀, N.D. Jago, Brit. Mus.1965-474 ( NHMUK 010926529);   25.v.[19]59, N.D. Jago, Brit. Mus.1965-474 ( NHMUK 010926579).  Western Reg.[ion] mls E of Elmina, 1°20W 5°05.5N,  xii.1960, 1♂, 1♀, N.D. Jago, identified  Xenogryllus eneopteroidesby P.C. Tinning, 1966, Brit. Mus.1965-474 ( NHMUK0 10926509, 010926511). T.v.T [Trans-Volta Togo], Ho-Hohoe rd. 10 milesfrom Voltabridge,  iii.1960, N.D. Jago, Brit. Mus.1965-474 ( NHMUK 010926553).  Western Reg. Asankragua-Enchi rd., TanoR. Ferry,  24.ix.1962, 1♂, N.D. Jago, Brit. Mus.1965-474 ( NHMUK 010926510).   Guinea: Kaoulenta[Kéoulenta], [Mount] Nimba( Guinée), ii–vi.[19]42, M. Lamotte, 1♂, 1♀, 2 juveniles( MNHN).  Nimba( Guinée), [Mount] Pierre Richaud,  900 m, M. Lamotteii–iv.[19]42, 1♀, identified  Xenogryllus eneopteroidesBol.by L. Chopard; 1♀( MNHN).  Nimba( Guinée), Lamotte, Amiet, Vanderplaetsen,  xii.56–  v.57: Ziéla,  17.ii.[19]57, 1 juvenile; Ziéla,  20.iv.[19]57, 1 juvenile; sav. [savannah] entre Ziéla et Gbakbré,  9.xii.1956, 1 juvenile; Zgpo.[Zouguépo],  900 m, androp,  18.iv.[1957], 2♀; sav. [savannah] Ziéla,  3.ii.[19]57, 1 juvenile; sav. [savannah] Ziéla, 20–25.xii.[19]56, 4 juveniles; sav. [savannah] Kéoulenta,  5.xii.[19]56, 2 juveniles; forêt Ziéla,  19.ii.[19]57, 1 juvenile; 1 juvenile, A.V. n°23, identified  Xenogryllus eneopteroidesBol.larva by L. Chopard; 1 juvenile, A.V. n°38; 1 juvenile, 87CD;  14.iii.[19]57, 1 juvenile, fge n°47 ( MNHN).  Mt. Nimba, Mt. Leclerc[Mount Jean Charles Leclerc],  1500 m,  23.iii.1991. M. Lamotte, n°319, 1♀( MNHN);  1♀, molecular sample  X22-XenGU (MNHN-EO-ENSIF1494). Nimba( Guinée), M. Lamotte,  xii.56–  v.57, 29.x.[1956], 1♂, herbes A2 Ziela, identified  Xenogryllus eneopteroidesBol.by R. Roy, 1959 ( MNHN).  Guinée Française, Diéké[Diecke], P. Chabamaud, 1920, 1♀, identified  Xenogryllus eneopteroidesBol.by L. Chopard( MNHN).   Ivory Coast: Lamto( Toumodi), Coll. E.N.S. Paris, Lamotte et collab.: 1–  10.ii.[19]62, 3♂, 2♀; 1963, 1♂, Xen-Robillardmorpho; 25–31.xi.[19]62, 1♀; 15–20.viii.[19]62, 1 juvenile; iv.[19]62, 1♀;  7.xi.[19]62, 1♀;  20.xi.[19]62, 1♀; vi.[19]62, 1♀;  4.vii.[19]62, 1♀, mare sav. inondée [inundated savannah] ( MNHN).  Forêtà 15 kmde Lamto,  10.ix.1968, C. Girard, 1♀, 1 juvenile( MNHN).  Forêt de Yape, 1♂( MNHN).  Toumodi,  xii.1930–  iv.1931, Ch. Alluaud& P. A. Chapuis, 1♂( MNHN).  Danané, i.1939, 1♀ ( MNHN).  Lamto( Toumodi), C. Girard:  12.ii.1968, savane [savannah], lisière forêt galerie [hedge of gallery forest] 6♂, 3♀, 5 juveniles; 1♂, identified  Xenogryllus eneopteroidesBol.by R. Roy, 1974;  12.iii.1968, lisière, forêt, savane, 1♂, molecular sample XenCI (MNHN-EO- ENSIF1481); 8♂, 2♀, 7 juveniles; 1♂, photos MEB [SEM photos] (MNHN-EO-ENSIF1518);  15–30.vii.1968, savane [savannah], lisière forêt galerie [hedge of gallery forest] 1♂, 2♀, 3 juveniles; 27–28.  ii.1968, 2♀, 1 juvenile, à la lumière; 10–20. iv.1968, 1♂, à la lumière ( MNHN).  Lamto( Toumodi), Paul Planquette: 1♀, PNB 18; 1 juvenile, PNB 13B;  15.i.[19]64, 1♀, PNB 64 ( MNHN).  Lamto( Toumodi),  30.vi.[19]62, Coll. E.N.S. Paris, Lamotte et al., 1♂, identified  Xenogryllus eneopteroidesby L. Chopard( MNHN).  Nimba[Mount]: Lamotte, 1946, Z4, 1♀, identified  Xenogryllus eneopteroidesby R. Roy, 1967 ( MNHN).  Lamto, 1963, 1♂( MNHN).  Assinie, 3295-85, Chaper, 1♂( MNHN).  Réserve du Banco[Banco national park], R. Paulian& G. Delamare, 2♂, 2♀, 2 juveniles( MNHN).  Bouaké(C.I.),  22.v.[19]62, coll. E.N.S. Paris, Lamotte et collab., 1♂( MNHN).  Taï,  18.viii.1978, G. Couturier, 1♀( MNHN).  Lamto, 1962, P. Le Gall, 2♂, 2♀( MNHN).   Kenya:CRS [Catholic Relief Service] Tebere, 0039.5S 3723E,  10.i.[19]72, I.A. D.R.72/58, 1♀, Brit. Mus.1982-71 ( NHMUK 010926573).   Nigeria: Lagos, Ikoyi,  viii. 1951, L. Bala, 1♂, identified  Xeneogryllus eneopteroidesby L. Chopard( MNHN). U.C. [University College] Ibadan,  5.ix.1952, Tephrosia, leaf, Teph-66 coll. G.H. Caswell, 1♀( MNHN).  Ibadan,  6.vi.[19]23, 1♂, 1605 ( NHMUK 010926581).  Calabar, Nigeria,  10.iv.[18]82, 10.30 A.M., W. edge of [?t] Kwaswamp, E. edge of UNICAL staff quarters; about 50 cmup on base of tangle of turns e.g.  Clitonia? rubiginosa, notes 72, 1♀, C.I.E. coll A.156 76 Presby Comm Inst Ent B.M.1985-1( NHMUK 010926537).   Replublicof Congo: Sibiti,  xi-1963, mission A. Descarpentrieset A. Villiers  1963–1964, 2♂, 1♀( MNHN).  Baie de Lagoa, envoi H. Deyrolle, 1♀( MNHN).  Mayumbe, Luki, L. Tiebers[?], 1♀, Musée du Congo( MNHN).  Odzala,  x.1963, 3♂, 3♀, 1 juvenile( MNHN).  Congo, 1963, 1♀( MNHN).  Ile[island] M’Bamou,  26.vii.1968, Fidèle, 1♀( MNHN).  Boma, Sundi, P. Rolin, 1♂( RBINS).  Banana, F. Busschodts, 4♀( RBINS).   Sierra Leone: Piste Bandankoro,  29.v.1963, savane de plaine ayant brûlé [burnt lowland savannah], mission ENS-IFAN aux Monts Loma, 1♂, identified  Xenogryllus eneopteroidesby R. Roy, 1992; 2♂, 3♀( MNHN).  Piste Keimadugu, savane [savannah],  520 m,  29.v.1963, mission ENS-IFAN aux Monts Loma, 2♂, 1♀( MNHN).  Firawa, savane [savannah],  1.vi.1963, mission ENS-IFAN aux Monts Loma, 1♂, 1♀( MNHN).  Njala, E. Hargreaves:  21.vii.[19]32, B.M.1938-149, identified  X. eneopteroidesBol.by L. Chopard, pres Imp. Inst. Ent. Brit. Mus.1965-474 ( NHMUK 010926552);   x.1936, 1♂, pres Imp. Inst. Ent. Brit. Mus.1965-474, 1♂, identified  Xenogryllus eneopteroidesby B. Uvarov( NHMUK 010926534);   23.v.[19]25, 1♂, identified  X. eneopteroidesBol.by L. Chopard, B. M. 1938-149 ( NHMUK 010926530);   xi.1935, 1♂, pres Imp. Inst. Ent. Brit. Mus.1936-252, 1♂, identified  Xenogryllus eneopteroidesby B. Uvarov( NHMUK 010926576).   Uganda: Entebbe,  20.xi.1970, W.J. Bailey, B.M.1977-246, identified  Xenogryllus eneopteroidesby B. C. Townsend, 1978 ( NHMUK 010926570).  Kisaru,  19.vii.1983, 1♂, H.B. Johnson, at light ( NHMUK 010926551).   Zambia: Lusaka, about fifty metres from the Great East Road in Chalimbana, Chongwe Distr.,  19.xii.2013, William van Niekerk, 1♂, identified  X. eneopteroidesby William van Niekerk[online photograph]. Kapiri,  13.xi.1913, 2♀, L. Charliers, Musée du Congo, identified  Xenogryllus eneopteroidesby L. Chopard, R. Det.N 2814 ( MRAC).    Typelocality. Angola, Duque de Bragança, Bayão(=Kalandula).   Distribution.Sub-Saharan Africa: Angola, Benin, Cameroun, Central African Republic, Democratic Republic of the Congo, Gabon, Ivory Coast, Kenya(one large female specimen with unclear identification), Nigeria, Republic of Congo, Sierra Leone, Uganda, Zambia.   Emended diagnosis.Species of average size, close to  X. mozambicus  n. sp.and  X. maniema  n. sp., from which it differs by face almost flat in lateral view (more rounded in other species), shorter male FWs (short apical field), and shape of lophi in male genitalia. Differing from  X. lamottei  n. sp.and Asian species by following characters: well-carinated lateral angles of dorsal disc of pronotum (also carinated in  X. mozambicusand  X. maniema), male genitalia and very short ovipositor.   Redescription.Species of average size ( Fig. 1A–C), coloration gray brown little contrasted. Eyes large, lateral, occupying almost half of head height in lateral view ( Fig. 3A–B). Face almost flat in lateral view, with typical whitish mask underlined by a black line below eyes and on mandibles. Pronotum dorsal disc strongly carinated laterally ( Fig. 3B), coloration dark brown or brown, most often with a median dark brown band extended laterally near anterior margin, forming a T shape; lateral lobes almost homogeneously dark brown. First article of antennae dark brown.  Male.FWs very wide, dark coloration anterior to 1A including angle of 1A ( Fig. 5A). File with 658 stridulatory teeth (n=2) on transverse part of 1A. Harp longer than wide. Cell c1 narrowed posteriorly, its fusion with cell b1 slightly shorter than its individualized part. Cell c2 large, d2 long and narrow. Mirror large, wider than long and little rounded, its inner angle forming an angle. Apical field short, wider than long, including four or five cell alignments.  Male genitalia( Figs 7A–C, 8A–B): Pseudepiphallic lophi as long as rest of pseudepiphallic sclerite, forming a long rectangle, sometimes slightly widened preapically; lophi with a narrow membranous inner margin setose basally ( Fig. 8A–B); their bases fused until mid-length; apex pointed and divergent, ended by a thin lamella without hook-like inner dorsal expansion (in dorsal view, anterior base of lamella suggests that lamella may partly correspond to modified inner dorsal expansion of other species); ventral blade of lophi without longitudinal wrinkles, but with a strong ventral transverse carina, as in  X. mozambicus  n. sp.Pseudepiphallic parameres with a strong rectangular ventral lobe and a basal membranous lobe. Rami strong, their apex forming wide convergent hooks. Ectophallic apodemes strong, not lamellate. Ectophallic lateral expansions, lateral sclerites of ectophallic fold and endophallic sclerite partly fused, forming a wide ventral sclerotized plate, trifid apically; endophallic apodemes made of wide lateral lamellas and a narrow dorsal crest.  Female:( Fig. 1C). Dorsal disc of pronotum almost rectangular, its posterior margin slightly bisinuate. FWs dorsal field with 7–10 longitudinal veins (m=8, n=10). Subgenital plate with a shallow apical indentation ( Fig. 9A). Ovipositor ( Fig. 10A) very short, about one third of FIII length.   FIGURE 1.(A–C)  Xenogryllus eneopteroides: male in dorsal (A) and lateral (B) views, female in dorsal view (C); (D–G)  Xenogryllus marmoratus: male in dorsal (D) and lateral (E) views, female in dorsal (F) and lateral (G) views; (H–I)  Xenogryllus lamottei  n. sp.: male holotype in dorsal (H) and lateral (I) views; (J–K)  Xenogryllus maniema  n. sp.: male paratype in dorsal (J) and lateral (K) views; (L–O)  Xenogryllus maichauensis: male in dorsal (L) and lateral (M) views, female in dorsal (N) and lateral (O) views. Scale bar: 1 cm.  Female genitalia.Copulatory papilla ( Fig. 11A–B) conical and narrow, its apex rounded; well-sclerotized except base and apex.  Life history traits.According to Chiffaud & Gillon (1984),  X. eneopteroidesis found in Ivory Coastat the end of the dry season, in parts of savannah protected from fires. Their study revealed that the species feeds only on leaves and flowers of dicotyledones which are specific to savannah.  Calling song.According to the sonogram showed in Desutter (1983), analyzed by Robillard & Desutter- Grandcolas (2004a, 2011), the calling song of  X. eneopteroidesis a short echeme made of five long syllables (syllable duration = 65 ms, period = 106.5 ms. Echeme duration = 611 ms, echeme period = 1062 ms. The dominant frequency is low (3.6 kHz) and corresponds to the first peak of the spectrum.  Measurements.See Table 2.   Taxonomic discussion.The material examined reveals numerous slight differences in male genitalia ( Fig. 8A– B) and size along the wide geographic distribution of the species. In particular, specimens from Congodiffer from the material from the western coast of Africa. However, these differences are not sufficiently clear or are based on too few individuals to delimit different species yet. More information and sampling will be necessary to improve the study  Xenogryllusin Africa. 2147526258 MNCN Angola Duque de Braganca Bayao 5 306 1 1 lectotype 2147526434 [580,659,1673,1698] MNHN de Zagnamado & de Ketou & d'Aquaquere & P. Ducorps Benin Bas-Dahomey 5 306 1 1 2147526184 MNHN Cameroon Conradt 5 306 1 1 2147526317 [509,1433,1745,1770] MNHN de Kribi & Dr. Gromier Cameroon Reg 5 306 1 1 2147526316 [151,1356,1781,1806] MNHN Eitel Cameroon Pipindi Kamerun 5 306 1 1 2147526415 1975-07-10 NHMUK N. D. Jago & B. C. Townsend Cameroon Yaounde Nkolbisson Agronomy Station 5 306 NHMUK 010926525 1 1 2147526416 1995-04-07 MNHN L. Desutter-Grandcolas Central African Republic N'Gotto Republique Centrafricaine 5 306 2 1 1 Lobaye 2147526423 1964-03-05 NHMUK Tyson Roberts & Brit. Mus. Democratic Republic of the Congo Congo Coquilhatville Tumba 5 306 2 1 1 2147526506 1938-12-06 RBINS H. J. Bredo & R. Mus. Nat. Belg. Congo Belge Lukafa 6 307 1 1 2147526417 1936-03-05 NHMUK, AMNH H. J. Bredo & Acacia & Niangara 3.6666667 Aru 1307 3.6666667 Belgian Congo 6 307 8 2 6 2147526421 MRAC Musee du Congo & L. Chopard & R. Det. Haut-Ituri Blommaert Faradje 6 307 1 1 2147526428 MRAC S. E. Katanga & R. P. Claquin & Musee du Congo & Chopard & Det. Ngaye 6 307 1 1 2147526326 MRAC R. P. Claquin & Musee du Congo & L. Chopard & Det. 6 307 2 1 1 2147526325 1925-06-01 MRAC Musee du Congo & L. Chopard & R. Det. Haut-Uele 6 307 1 1 2147526427 MRAC Ch. Scops & Musee du Congo & L. Chopard & R. Det. Geti Kibali-Ituri 6 307 1 1 2147526257 1994-06-19 MNHN L. Desutter-Grandcolas Gabon Ekalla Plateaux Bateke 6 307 2 1 1 2147526502 MNHN L. Chopard Gabon Bas Ogooue 6 307 1 1 2147526190 [269,1160,620,645] 19. vii. RBINS A. Pauly Gabon Ntoum 6 307 6 2 2 2 2147526424 1959-12-23 1963-07-19 1959-12-23 NHMUK Kade A. R. S. & I. K. Bacheampong & N. D. Jago & Brit. Mus. Ghana Hansowodze 6 307 NHMUK 010926535 5 5 Eastern 2147526425 1959-05-03 1963-07-11 1959-05-03 NHMUK I. K. Bacheampong & N. D. Jago & Brit. Mus. Ghana Brit. Mus. Kade Agr. Res. Stn 6 307 NHMUK 010926529 3 1 2 Eastern 2147526259 [662,1433,836,861] 25. v. NHMUK N. D. Jago & Brit. Mus. Ghana 6 307 NHMUK 010926579 1 Eastern 2147526323 1960-03 1960-12-01 1960-03 NHMUK N. D. Jago & P. C. Tinning & Brit. Mus. Ghana 5.0916667 Ho-Hohoe rd. Western Reg. 6 307 NHMUK 010926553 1 1 Volta 2147526188 1962-09-24 NHMUK N. D. Jago & Brit. Mus. Ghana Western Reg. Asankragua-Enchi rd. Ferry Tano 6 307 NHMUK 010926510 1 1 Volta 2147526255 MNHN M. Lamotte Guinea-Bissau Kaoulenta Guinee Nimba 6 307 4 1 2 1 2147526429 MNHN M. Lamotte & L. Chopard Guinea-Bissau Nimba 900 Pierre Richaud Guinee 6 307 2 2 2147526420 1956-12 1957-05 1956-12 MNHN L. Chopard Guinea-Bissau Nimba 900 Lamotte Guinee 6 307 17 2 15 2147526261 1991-03-23 MNHN M. Lamotte Guinea-Bissau 1500 Mt. Leclerc Mt. Nimba 6 307 1 1 2147526318 1922-10 1957-05-29 1922-10 MNHN Nimba & R. Roy Guinea-Bissau Guinee Ziela Lamotte 6 307 2 1 1 2147526433 MNHN P. Chabamaud & L. Chopard Guinea-Bissau Dieke Guinee Francaise 6 307 1 1 2147526439 10. ii. 4. vii. 10. ii. MNHN Coll. E. N. S. & Paris Ivory Coast Lamto Xen-Robillardmorpho Toumodi 6 307 13 8 1 4 2147526280 1968-09-10 MNHN C. Girard Ivory Coast Lamto Foret 6 307 2 1 1 2147526437 [719,1056,1556,1581] MNHN Ivory Coast Foret de Yape 6 307 1 1 2147526438 1930-12 1931-04 1930-12 MNHN Ch. Alluaud & P. A. Chapuis Ivory Coast Toumodi 6 307 1 1 2147526322 [619,960,1591,1616] MNHN Ivory Coast Danane 6 307 1 1 2147526321 1968-02-02 1968-07-30 1968-02-02 MNHN Ivory Coast Lamto Girard Toumodi 6 307 44 9 16 19 2147526185 15. i. MNHN Ivory Coast Lamto Paul Planquette Toumodi 6 307 3 2 1 2147526432 30. vi. MNHN Coll. E. N. S. & Paris & L. Chopard Ivory Coast Toumodi Lamto 6 307 1 1 2147526197 MNHN R. Roy Ivory Coast Lamotte Nimba 6 307 1 1 2147526186 [1012,1331,1879,1904] MNHN Ivory Coast Lamto 6 307 1 1 2147526327 MNHN Chaper Ivory Coast Assinie 6 307 1 1 2147526194 MNHN R. Paulian & G. Delamare Ivory Coast Reserve du Banco 6 307 6 2 2 2 2147526193 [396,1374,1952,1977] 22. v. MNHN E. N. S. Paris Ivory Coast Bouake 6 307 1 1 2147526501 1978-08-18 MNHN G. Couturier Ivory Coast Tai 6 307 1 1 2147526196 [611,1092,1988,2013] MNHN P. Le Gall Ivory Coast Lamto 6 307 4 2 2 2147526418 10. i. NHMUK D.R. Ragge & Brit. Mus. Kenya 7 308 Tebere 6 307 NHMUK 010926573 1 1 2147526268 [151,1433,188,213] 1951-08 MNHN L. Bala & Xeneogryllus & L. Chopard Nigeria Ikoyi 7 308 1 1 Lagos 2147526285 [151,1340,223,248] 1952-09-05 MNHN Tephrosia & G. H. Caswell Nigeria Ibadan 7 308 1 1 Lagos 2147526271 6. vi. NHMUK Nigeria Ibadan 7 308 NHMUK 010926581 1 1 Lagos 2147526328 10. iv. NHMUK Kwa & Clitonia & Pres & Comm Inst Ent B. M. Nigeria Nigeria Calabar 7 308 NHMUK 010926537 1 1 Lagos 2147526430 1963 1964 1963 MNHN A. Descarpentries & A. Villiers Democratic Republic of the Congo Sibiti Replublic 7 308 3 1 2 2147526284 MNHN H. Deyrolle Democratic Republic of the Congo Baie de Lagoa 7 308 1 1 2147526329 [528,1248,404,429] MNHN Luki, L & Musee du Congo Democratic Republic of the Congo Tiebers Mayumbe 7 308 1 1 2147526256 1963-10-03 MNHN Democratic Republic of the Congo Odzala 7 308 7 3 1 3 2147526263 [484,791,439,464] MNHN Democratic Republic of the Congo Congo 7 308 1 1 2147526288 [801,1433,439,464] 1968-07-26 MNHN Fidele Democratic Republic of the Congo M'Bamou Ile 7 308 1 1 2147526276 [151,583,476,501] RBINS Sundi, P & Rolin Democratic Republic of the Congo Boma 7 308 1 1 2147526508 [599,1031,476,501] RBINS Banana, F Democratic Republic of the Congo Busschodts 7 308 4 4 2147526273 1963-05-29 MNHN Monts Loma & R. Roy Sierra Leone Piste Bandankoro 7 308 6 3 3 2147526192 1963-05-29 MNHN Monts Loma Sierra Leone 520 Piste Keimadugu 7 308 3 1 2 2147526319 1963-06-01 MNHN Monts Loma Sierra Leone Firawa 7 308 2 1 1 2147526324 21. vii. NHMUK Blommaert, Musee du Congo & L. Chopard & Imp. Inst. Ent. Brit. Mus. Sierra Leone Hargreaves Njala 7 308 NHMUK 010926552 1 2147526422 1936-10-01 NHMUK Imp. Inst. Ent. Brit. Mus. & B. Uvarov Sierra Leone 7 308 NHMUK 010926534 2 2 2147526264 23. v. NHMUK Chopard Sierra Leone 7 308 NHMUK 010926530 1 1 2147526274 1935-11-01 NHMUK Imp. Inst. Ent. Brit. Mus. & B. Uvarov Sierra Leone 7 308 NHMUK 010926576 2 2 2147526200 1970-11-20 NHMUK W. J. Bailey & B. C. Townsend Uganda Entebbe 7 308 NHMUK 010926570 1 2147526267 1983-07-19 NHMUK H. B. Johnson Uganda Kisaru 7 308 NHMUK 010926551 1 1 2147526320 1913-11-13 2013-12-19 1913-11-13 MRAC William van Niekerk & L. Charliers & Musee du Congo & L. Chopard & R. Det. Zambia Great East Road in Chalimbana Kapiri Chongwe Distr. 7 308 3 2 1 Lusaka 2147526187 [199,927,1015,1040] Angola Duque de Braganca Bayao 7 308 1 holotype