Revision of the southern African genus Frankenbergerius Balthasar with description of new taxa (Coleoptera: Scarabaeidae: Scarabaeinae) Frolov, A. V. Scholtz, C. H. Journal of Natural History 2005 2005-06-29 39 25 2355 2377 5HDD6 Frolov & Scholtz, 2005 Frolov & Scholtz 2005 [391,916,1304,1328] Insecta Scarabaeidae Frankenbergerius Animalia Coleoptera 18 2371 Arthropoda subSpecies armatus subsp. nov. tuberculatus  ( Figures 17–21, 24, 26, 27)  Diagnosis This subspecies differs from the nominotypical one chiefly by the shape of the clypeal horns in males: the horns curve upwards, with acute apices ( Figures 17, 21).  Description  Holotype: „( Figure 21). Body length 9.5 mm, width 5.5 mm. Dorsal surface of pronotum, elytra and head with minute setae.  Head.Clypeus with two horns slightly curved upwards; horns acute at apices. Genae obtuse, finely bordered. Genal sutures visible as fine lines. Lateral margin of clypeus not or very feebly sinuate near genal suture. Frontal suture broadly interrupted at middle. Area behind genae slightly concave, coarsely punctate with big adjoining punctures with distinct margins. Genae and anterior part of clypeus densely punctate; margins of punctures indistinct. Very anterior part of clypeus sculpture nearly rugose. Disc of head with minute, feebly visible punctuation. Inner (dorsal) surface of clypeal horns finely granular; outer (ventral) surface smooth. Anterior margin of clypeus flattened, deeply sinuate at middle, slightly rugose.  Pronotum.Anterior angles rounded. Lateral margins with fine border, hind and anterior angles and anterior margin with wider border. Border of anterior margin widened and sinuate at middle. Base not bordered. Hind angles obtuse. Dorsal surface densely and coarsely punctate; punctures separated by 0.2–0.3 times their diameters on disc, becoming denser laterally.  Elytra.Striae fine, becoming slightly wider and deeper apically, shiny, punctate (punctures separated by three to four puncture diameters). Intervals flat on disc, feebly convex in apical part, smooth to slightly shagreened and punctate (punctures separated by 1.5 to two puncture diameters). Intervals 2–7 with longitudinal tubercles more developed than those in  F. armatus armatus. Interval 8 with long keel interrupted at middle of elytron. Interval 9 with smaller keel situated at about middle of elytron. Keels, especially hind part of keel on interval 8, serrate in most individuals.  Underside.Pygidium with deep border, disc densely punctate. Abdominal and thoracic sternites coarsely punctate except for disc of meso- and metasternum.  Legs.Anterior tibiae with smaller teeth between major outer teeth in most specimens. Spur of anterior tibia bifurcated in males and simple, acute in females.  Aedeagus.Similar to  F. armatus armatus( Figure 17).  Paratypes: body length of males 7.0–11.0 mm, width 4.1–5.2 mm, of females 6.3–8.6 mmand 4.0– 5.1 mm. Colour of body from black to dark brown. In most specimens, disc of head and pronotum without setae. Length of clypeal horns varies in males ( Figures 18, 19). Females can be separated by acute, not bifurcated spur of anterior tibiae and by shorter clypeal processes in some cases ( Figure 20).  Distribution This subspecies is distributed is Mpumalanga and Limpopoprovinces. The southernmost known locality is Wakkerstroom in southern Mpumalanga( Figure 27).  Type material   Limpopo Province: Entabeni Forest Station[ 23 °00 9S, 30 °14 9E],  November 1931, 3♀, G. van Sonleg. ( TMSA). Malta[ 24 °10 9S, 30 °14 9E],  1 February 1927, 1♀( TMSA); Uitzicht [ 23 °19 9S, 30 °01 9E], Soutpansberg Distr.,  November 1924, 1 „and 1♀( IRSNB); Brak River [? 22 °36 9S, 29 °44 9E],  October 1927, 1 „( TMSA). Mpumalanga: Nelshoogte, Knuckles Rocks Forest, 25 °47 9S, 30 °50 9E,  24 October 1986, intercept traps, 41 day, 6 „ 3♀, S. Endrödy-Youngaleg. ( TMSA). Wakkerstroom [ 27 °21 9S, 30 °08 9E],  January 1925, 1♀, G. van Damleg. ( TMSA). Uitsoek, 25 °15 9S, 30 °34 9E,  7 February 1987, high altitude grassveld, pitfall traps, 61 days, 9 „and 7♀;  5 December 1986, Grootkloof indigenous forest, 25 °15 9S, 30 °33 9E, pitfall traps, 53 days, 5 „and 6♀, S. Endrödy-Youngaleg. ( TMSA). Nelshoogte,  11 February 1987, Knuckles grassveld, Agaricaceaefungus, 1 „; pitfall traps, 58 days, 2 „and 4♀;  4 December 1986, Knuckles Rocks Forest, pitfall traps, 68 days, 2 „;  8 April 1987, Knuckles grassveld, grass-netting, 1♀, S. Endrödy-Youngaleg. ( TMSA). Berlin, 24 °32 9S, 30 °44 9E,  4 February 1987, pitfall traps, 41 days, 1 „? and 1♀, S. Endrödy-Youngaleg. ( TMSA). Mariepskop, 24 °35 9S, 30 °50 9E,  2 May 1981, pitfall traps, 5 days, 1 „, S. Endrödy-Youngaleg. ( TMSA). Road Barbeton to Bulembu near Swazilandborder [ 25 °55 9S, 31 °10 9E],  20 January 2000, 1 „and 1♀( TMSA). White River Distr., farm Lichtfontein [ 25 °19 9S, 31 °01 9E],  28 December 1992, 1 „, R. Müllerleg. ( TMSA). 1924-11 2000-01-20 1924-11 TMSA, IRSNB G. van Son & S. Endrody-Younga & G. van Dam & R. Muller Malta Knuckles Rocks Forest 20 2373 -25.919167 Limpopo Province 20 31.169167 Soutpansberg Distr. 19 2372 29 29