Paulian, 1977 : 1215
Paulian, 1977 : 1219
Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood
Frolov, Andrey V.
Montreuil, Olivier
Akhmetova, Lilia A.
Zootaxa
2016
4207
1
1
93
Paulian, 1977
Paulian
1977
[151,557,692,718]
Insecta
Orphnidae
Triodontus
Animalia
Coleoptera
79
80
Arthropoda
species
alticola
Triodontus alticola Paulian, 1977: 1215. Triodontus maroantsetrae Paulian, 1977: 1219, new synonym
Typematerial examined. Triodontus alticola. Holotype( Figs. 41C, D), male, “ MadagascarAmbohitsitondrona [ Toamasina Province] XI-1949 Michel/ Triodontus alticola n. sp. R. Pauliandet. / TYPE” ( MNHN). Paratypes: two females with the same locality and date label ( MNHN); two males and two females with the same locality label but the date “ XII-52” ( MNHN); one male, “ MadagascarSambava [ Antsiranana Province] XI [...] Vadon!” ( MNHN); one male, “ MadagascarAmbodivoangy [ Toamasina Province] I.1950 Vadon!” ( MNHN). Triodontus maroantsetrae. Holotype, male, “ MadagascarRon Maroantsetra XII.35 Vadon Antakotako/ MUSEUM PARIS1938 J. Vadonet E. Lebis/ TYPE/ Triodontus maroantsetrae n. sp. R. Pauliandet.” ( MNHN). Additional material examined. MADAGASCAR: Toamasina:two males, “ Madagascar Ambohitsitondrona II-1948 Michel[ 15°34′59″S, 50°01′00″E]” ( MNHN); one male, “ MadagascarAmbodivoangy [ 15°17′49″S, 49°36′47″E] I.1946Vadon!” ( MNHN); one male, same label but XII.1947(MNHN); one female, “ MadagascarAntakotako [ 15°19′00″S, 49°48′00″E] II.1946Vadon!” ( MNHN); one male, “ MadagascarEst distr. Mananara - N. Antanambe Vadonet Peyrieras [ Mananara District, Antanambe, 16°25′59″S, 49°51′00″E]” ( MNHN); four males, one female, “ MadagascarAmbohitsitondrona I-1946 Vadon! [ 15°34′59″S, 50°01′00″E]” ( MNHN); one male, “ MadagascarAmbohitsitondrona 25-I-1948Michel [ 15°34′59″S, 50°01′00″E, 25.I.1948]” ( MNHN); one male, one female, “ MadagascarAmbohitsitondrona X-1947 Vadon! [ 15°34′59″S, 50°01′00″E]” ( MNHN).
Diagnosis. Triodontus alticolacan be separated from other species of the genus by the characteristic shape of the parameres having the outer lobes long, curved downward and inward; with more-or-less separated, strongly sclerotized lateral parts and membranous, overlapping medial parts; and without lateral notches.
Description. Male.Color of head, pronotum, and elytra dark brown, legs, antennae, and underside of the body slightly paler ( Figs. 41A, C). Frontoclypeus slightly convex anteriorly, obtusely rounded laterally, anterior margin setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small, dorsal and large, ventral parts. Frontoclypeus with tubercles mediad of each eye and with a horn in center. The horn is longer than width of the head, acutely rounded apically, somewhat curved caudally, with slightly flattened anterior side and slightly rugose posterior side. Pronotum with a bulge medially in center, with excavations each side of bulge, and with 2 ridge-shaped tubercles laterally of each excavation. The median bulge with 2 distinct tubercles and somewhat excavated between the tubercles. Lateral margins with wide border bearing a row of brown setae. Anterior margin with wide, smooth border, almost not sinuate medially. Posterior margin with fine border, not crenulate, punctate with a few narrow, longitudinal punctures. Surface of most pronotum punctate with minute, feebly visible puctures. Scutellum rounded apically, visible part is about 1/15 length of elytra. FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H). Elytra convex, with distinct humeral humps. Maximum width approximately at middle. Elytra with 10 striae; stria 1 (sutural) distinct, other feebly marked with small punctures. Intervals with minute punctures, almost smooth. Base of elytra with an irregular row of coarse punctures. Abdominal sternite 8 medially deeply sinuate, without tubercles or concavities in middle ( Fig. 41G). Parameres ( Fig. 41E) with outer lobes long, curved downward and inward; with more-or-less separated, strongly sclerotized lateral parts and membranous, overlapping medial parts; and without lateral notches. Internal sac of aedeagus with a hoof-shaped sclerite ( Fig. 41F). Female.Females ( Fig. 41B) differ from males in having a protibial apical spur; absence of head and pronotum armature including tubercles mediad of each eye; and longer abdominal sternites including 6th sternite, which is evenly rounded apically. Variation.The pronotal and head armature of males is less developed in some specimens. Body length of examined specimens varied from 8.8–12.0 mm (males) and from 9.0–11.0 mm (females).
Distribution.This species is known from a few localities in the northeastern part of the island, mostly north of Antongil Bay ( Fig. 41H).
Remarks. Triodontus maroantsetraewas described from one male specimen from Antakotako. Comparison of this specimen with the typeseries of T. alticolahas shown that they have similar shape of the parameres. The only difference is the feebly developed frontoclypeal horn and almost absent ridges and tubercles on the pronotum in the holotypeof T. maroantsetrae. We believe all these specimens fall within the limits of allometric variability of one species, therefore the new synonymy is here proposed.
1416610143
1949-11
TYPE, MNHN
Michel
Madagascar
R. Paulian
Toamasina Province
79
80
3
Toamasina Province
holotype
1416610150
[199,1009,967,992]
MNHN
79
80
1
paratype
1416610139
1952-12
MNHN
79
80
1
paratype
1416610105
MNHN
Madagascar
Vadon
79
80
1
Antsiranana Province
paratype
1416610104
[417,1409,1039,1064]
1950-01
MNHN
Madagascar
Vadon
79
80
1
Toamasina Province
paratype
1416610145
1935-12
MUSEUM, PARIS, TYPE, MNHN
J. Vadon & E. Lebis
Madagascar
R. Paulian
Vadon Antakotako
79
80
PARIS 1938
2
holotype
1416610141
1948-02
MNHN
Michel
Madagascar
-15.5830555
Madagascar Ambohitsitondrona
21
50.016666
79
80
1
Toamasina
1416610144
1946-01
MNHN
Madagascar
-15.296945
Madagascar
21
49.613052
79
80
1
1416610134
1946-02
MNHN
Madagascar
-15.316667
Madagascar
21
49.8
79
80
1
1416610128
MNHN
N. Antanambe Vadon
Madagascar
-16.433054
Mananara District
21
49.85
79
80
1
1416610140
1946-01
MNHN
Vadon
Madagascar
-15.5830555
Madagascar
21
50.016666
79
80
1
1416610126
1948-01-25
MNHN
Madagascar
-15.5830555
Madagascar
21
50.016666
79
80
1
1416610154
1947-10
MNHN
Vadon
Madagascar
-15.5830555
Madagascar
21
50.016666
79
80
1