Paulian, 1977 : 1215 Paulian, 1977 : 1219 Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood Frolov, Andrey V. Montreuil, Olivier Akhmetova, Lilia A. Zootaxa 2016 4207 1 1 93 Paulian, 1977 Paulian 1977 [151,557,692,718] Insecta Orphnidae Triodontus Animalia Coleoptera 79 80 Arthropoda species alticola     Triodontus alticola  Paulian, 1977: 1215.    Triodontus maroantsetrae  Paulian, 1977: 1219,  new synonym     Typematerial examined.  Triodontus alticola. Holotype( Figs. 41C, D), male, “ MadagascarAmbohitsitondrona [ Toamasina Province]  XI-1949 Michel/  Triodontus alticola n. sp. R. Pauliandet. /  TYPE” ( MNHN).   Paratypes: two females with the same locality and date label ( MNHN); two males and two females with the same locality label but the date “  XII-52” ( MNHN); one male, “ MadagascarSambava [ Antsiranana Province] XI [...] Vadon!” ( MNHN); one male, “ MadagascarAmbodivoangy [ Toamasina Province]  I.1950 Vadon!” ( MNHN).    Triodontus maroantsetrae. Holotype, male, “ MadagascarRon Maroantsetra  XII.35 Vadon Antakotako/ MUSEUM  PARIS1938 J. Vadonet E. Lebis/  TYPE/ Triodontus  maroantsetrae n. sp. R. Pauliandet.” ( MNHN).   Additional material examined. MADAGASCAR: Toamasina:two males, “ Madagascar Ambohitsitondrona  II-1948 Michel[ 15°34′59″S, 50°01′00″E]” ( MNHN); one male, “ MadagascarAmbodivoangy [ 15°17′49″S, 49°36′47″E]  I.1946Vadon!” ( MNHN); one male, same label but XII.1947(MNHN); one female, “ MadagascarAntakotako [ 15°19′00″S, 49°48′00″E]  II.1946Vadon!” ( MNHN); one male, “ MadagascarEst distr. Mananara - N. Antanambe Vadonet Peyrieras [ Mananara District, Antanambe, 16°25′59″S, 49°51′00″E]” ( MNHN); four males, one female, “ MadagascarAmbohitsitondrona  I-1946 Vadon! [ 15°34′59″S, 50°01′00″E]” ( MNHN); one male, “ MadagascarAmbohitsitondrona  25-I-1948Michel [ 15°34′59″S, 50°01′00″E,  25.I.1948]” ( MNHN); one male, one female, “ MadagascarAmbohitsitondrona  X-1947 Vadon! [ 15°34′59″S, 50°01′00″E]” ( MNHN).   Diagnosis.  Triodontus alticolacan be separated from other species of the genus by the characteristic shape of the parameres having the outer lobes long, curved downward and inward; with more-or-less separated, strongly sclerotized lateral parts and membranous, overlapping medial parts; and without lateral notches.   Description. Male.Color of head, pronotum, and elytra dark brown, legs, antennae, and underside of the body slightly paler ( Figs. 41A, C). Frontoclypeus slightly convex anteriorly, obtusely rounded laterally, anterior margin setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small, dorsal and large, ventral parts. Frontoclypeus with tubercles mediad of each eye and with a horn in center. The horn is longer than width of the head, acutely rounded apically, somewhat curved caudally, with slightly flattened anterior side and slightly rugose posterior side. Pronotum with a bulge medially in center, with excavations each side of bulge, and with 2 ridge-shaped tubercles laterally of each excavation. The median bulge with 2 distinct tubercles and somewhat excavated between the tubercles. Lateral margins with wide border bearing a row of brown setae. Anterior margin with wide, smooth border, almost not sinuate medially. Posterior margin with fine border, not crenulate, punctate with a few narrow, longitudinal punctures. Surface of most pronotum punctate with minute, feebly visible puctures. Scutellum rounded apically, visible part is about 1/15 length of elytra.   FIGURE 41.  Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H). Elytra convex, with distinct humeral humps. Maximum width approximately at middle. Elytra with 10 striae; stria 1 (sutural) distinct, other feebly marked with small punctures. Intervals with minute punctures, almost smooth. Base of elytra with an irregular row of coarse punctures. Abdominal sternite 8 medially deeply sinuate, without tubercles or concavities in middle ( Fig. 41G). Parameres ( Fig. 41E) with outer lobes long, curved downward and inward; with more-or-less separated, strongly sclerotized lateral parts and membranous, overlapping medial parts; and without lateral notches. Internal sac of aedeagus with a hoof-shaped sclerite ( Fig. 41F).  Female.Females ( Fig. 41B) differ from males in having a protibial apical spur; absence of head and pronotum armature including tubercles mediad of each eye; and longer abdominal sternites including 6th sternite, which is evenly rounded apically.  Variation.The pronotal and head armature of males is less developed in some specimens. Body length of examined specimens varied from 8.8–12.0 mm (males) and from 9.0–11.0 mm (females).   Distribution.This species is known from a few localities in the northeastern part of the island, mostly north of Antongil Bay ( Fig. 41H).   Remarks.  Triodontus maroantsetraewas described from one male specimen from Antakotako. Comparison of this specimen with the typeseries of  T. alticolahas shown that they have similar shape of the parameres. The only difference is the feebly developed frontoclypeal horn and almost absent ridges and tubercles on the pronotum in the holotypeof  T. maroantsetrae. We believe all these specimens fall within the limits of allometric variability of one species, therefore the new synonymy is here proposed. 1416610143 1949-11 TYPE, MNHN Michel Madagascar R. Paulian Toamasina Province 79 80 3 Toamasina Province holotype 1416610150 [199,1009,967,992] MNHN 79 80 1 paratype 1416610139 1952-12 MNHN 79 80 1 paratype 1416610105 MNHN Madagascar Vadon 79 80 1 Antsiranana Province paratype 1416610104 [417,1409,1039,1064] 1950-01 MNHN Madagascar Vadon 79 80 1 Toamasina Province paratype 1416610145 1935-12 MUSEUM, PARIS, TYPE, MNHN J. Vadon & E. Lebis Madagascar R. Paulian Vadon Antakotako 79 80 PARIS 1938 2 holotype 1416610141 1948-02 MNHN Michel Madagascar -15.5830555 Madagascar Ambohitsitondrona 21 50.016666 79 80 1 Toamasina 1416610144 1946-01 MNHN Madagascar -15.296945 Madagascar 21 49.613052 79 80 1 1416610134 1946-02 MNHN Madagascar -15.316667 Madagascar 21 49.8 79 80 1 1416610128 MNHN N. Antanambe Vadon Madagascar -16.433054 Mananara District 21 49.85 79 80 1 1416610140 1946-01 MNHN Vadon Madagascar -15.5830555 Madagascar 21 50.016666 79 80 1 1416610126 1948-01-25 MNHN Madagascar -15.5830555 Madagascar 21 50.016666 79 80 1 1416610154 1947-10 MNHN Vadon Madagascar -15.5830555 Madagascar 21 50.016666 79 80 1