Grapsus ( Pachysoma ) quadratus – De Haan, 1835: 62 Sesarma dehaani H. Milne Edwards, 1853: 184 Stimpson, 1858: 106 Man, 1887: 642 Bürger, 1893: 615 Ortmann, 1894: 718 Stimpson, 1907: 134 Yamaguchi & Baba , 1993: 478 Fransen et al., 1997: 128 Wang & Leung, 2001: 31 Sesarma hanseni Rathbun, 1897: 92 Abele, 1975: 48 Sesarma ( Holometopus ) dehaani Tesch, 1917: 158 Balss, 1922: 154 Sakai, 1936: 234 Sakai, 1939: 682 Shen, 1940a: 97 Kamita, 1941: 217 Sakai, 1965: 202 Serène, 1968: 107 Baba & Miyata, 1971: 54 Kim, 1973: 487 Sakai, 1976: 655 Matsuzawa, 1977 Dai et al., 1986: 488 Nagai & Nomura, 1988: 43 Dai & Yang, 1991: 534 Huang, 1994: 597 Muraoka, 1998: 53 Holometopus dehaani Takeda, 1975: 147 Soh, 1978: 9 Takeda, 1982: 220 Chiromantes haematocheir C. haematocheir Orisarma dehaani O. neglectum O. patshuni O. patshuni Chiromantes dehaani Miyake, 1983: 179 Ng, 1999: 229 Minemizu, 2000: 298 Kobayashi, 2000: 122 Ng et al., 2001: 41 Kwok & Tang, 2005: 3 Takeda & Ueshima, 2006: 93 Lee, 2008: 135 Yang et al., 2008: 801 Naderloo & Schubart, 2009: 61–63 Hong et al., 2010: 259 Liu & Wang, 2010: 57 Huang et al., 2011: 732 Li & Chiu, 2013: 40 Komai & Ng, 2013: 540 Toyota & Seki, 2014: 190 Ng et al., 2017b: 102 “ Chiromantes ” dehaani Ng et al., 2008a: 220 Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species Schubart, Christoph D. Ng, Peter K. L. Raffles Bulletin of Zoology 2020 2020-12-23 68 891 994 84H2J (H. Milne Edwards, 1853) H. Milne Edwards 1853 [876,1388,1437,1461] Malacostraca Sesarmidae Orisarma Animalia Decapoda 13 904 Arthropoda species dehaani  ( Figs. 1C, D, 3B, 4A, B, D, F, 5D–F, 7E–G, 9C, 11A–G, 43B, 53A–E)      Grapsus( Pachysoma) quadratus– De Haan, 1835: 62, pl. 8 fig. 3.     Sesarma dehaaniH. Milne Edwards, 1853: 184;  Stimpson, 1858: 106(part); De Man, 1887: 642;  Bürger, 1893: 615;  Ortmann, 1894: 718;  Stimpson, 1907: 134(part);   Yamaguchi& Baba, 1993: 478, fig. 18b;  Fransen et al., 1997: 128;  Wang & Leung, 2001: 31, unnumbered figure.     Sesarma hanseni Rathbun, 1897: 92;  Abele, 1975: 48, figs. 1, 2.    Sesarma( Holometopus) dehaani–  Tesch, 1917: 158;  Balss, 1922: 154(part);  Sakai, 1936: 234, pl. 65, fig. 1;  Sakai, 1939: 682, pl. 77, fig. 1;  Shen, 1940a: 97;  Kamita, 1941: 217, text-fig. 120;  Sakai, 1965: 202, pl. 97, fig. 2;  Serène, 1968: 107;   Baba& Miyata, 1971: 54;  Kim, 1973: 487, text-fig. 217, pl. 95 fig. 167a, b, pl. 96 fig. 167c, d;  Sakai, 1976: 655, pl. 224 fig. 2;  Matsuzawa, 1977: pl. 110 fig. 2;  Dai et al., 1986: 488, text-fig. 274(2), pl. 68(7);  Nagai & Nomura, 1988: 43;  Dai & Yang, 1991: 534, text-fig. 274(2), pl. 68(7);  Huang, 1994: 597;  Muraoka, 1998: 53.    Holometopus dehaani–  Takeda, 1975: 147;  Soh, 1978: 9, 10, pl. 2d;  Takeda, 1982: 220, fig. 651.  Fig. 5. Chela. A, B,  Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; C,  C. haematocheir, male (18.6 × 20.7 mm) (RUMF-ZC-544), Japan; D–F,  Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; G,  O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; H,  O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong; I,  O. patshuni, male (20.4 × 18.2 mm) (ZRC 2012.0032), Hong Kong. A, D, G, H, I, outer view; B, C, F, dorso-lateral view; E, inner view.    Chiromantes dehaani–  Miyake, 1983: 179, pl. 60-2; Liu &  Ng, 1999: 229;  Minemizu, 2000: 298;  Kobayashi, 2000: 122, fig. 2n;  Ng et al., 2001: 41[includes references for this species from Taiwanup to 2001];  Kwok & Tang, 2005: 3, fig. 7;  Takeda & Ueshima, 2006: 93;  Lee, 2008: 135;  Yang et al., 2008: 801;  Naderloo & Schubart, 2009: 61–63, 67;  Hong et al., 2010: 259;  Liu & Wang, 2010: 57;  Huang et al., 2011: 732;  Li & Chiu, 2013: 40;  Komai & Ng, 2013: 540, figs. 1B, 6A, 7A–C, G;  Toyota & Seki, 2014: 190;  Ng et al., 2017b: 102[for other local references for Taiwan].   “ Chiromantes” dehaani–  Ng et al., 2008a: 220.   Material examined.  Lectotypemale (39.5 × 35.7 mm) ( RMNH D 157), Japan, coll. v. Siebold. Others: JAPAN– 1 male(35.0 × 38.5 mm) ( CBM-ZC 165), Obitsu Riverestuary, Kisarazu, Chiba Prefecture,  July 1990, coll. T. Furota; 1 male(31.0 × 28.3 mm) ( CBM-ZC 169), Hanami River, Chiba, Chiba Prefecture, coll. Nakano,  5 October 1990;  1 male(28.3 × 25.4 mm) ( CBM-ZC 11406), Yahata Channel, Ichihara, Chiba Prefecture, coll. J. Takayama,  30 August 2012; 6 males(23.9 × 22.0 mm to 42.0 × 37.7 mm),  3 females(23.9 × 22.0 mm to 29.5 × 26.9 mm) ( CBM-ZC 11458), Kaneda, Kisarazu, coll. T. Komai,  May 2009;  1 male(27.3 × 24.5 mm) ( ZRC1968.4.22.12), coll. T. Sakai, 1968;  1 female(24.3 × 21.5 mm) ( ZRC1970.8.27.1), Kamakura, Kanagawa Prefecture, Sagami Bay, coll. T. Sakai, 1968;  2 males(22.0 × 20.2 mm, 23.5 × 21.1 mm) ( CBM-ZC 7576), Tatara River, Fukuoka, Fukuoka Prefecture, coll. Y. Matsuzawa,  March 2003; 1 ovigerous female (21.4 × 19.4 mm) ( ZRC 2002.0226), river mouth, Izaku River, Satsuma Peninsula, Kagoshima Prefecture, coll. H. Suzuki,  30 August 2000;  2 males(larger 23.4 × 20.8 mm), 2 females(larger 25.2 × 22.0 mm) ( ZRC 2012.0057), Koutsuki River, Kagoshima City, coll. P.K.L. Nget al.,  24 October 2011;  1 male(25.6 × 23.0 mm), 1 ovigerous female (27.4 × 23.9 mm) ( CBM-ZC 3129), Tomari, Kume Island, Ryukyu Islands, coll. T. Komai,  12 June 1995;  1 male(38.3 × 36.6 mm) ( ZRC 2011.1026),  Zenda ForestPark, Kume Island, Ryukyu Islands, coll. T. Naruseet al.,  15 November 2009;  3 males(largest 38.4 × 35.5 mm), 1 female(26.6 × 23.7 mm) ( ZRC 2011.1027),  Zenda ForestPark, Kume Island, Ryukyu Islands, coll. P.K.L. Nget al.,  17 November 2006;  1 female(36.2 × 31.7 mm) ( CBM-ZC 9870), Ohtomi, Iriomote Island, Yaeyama Islands, coll. N. Shikatani,  29 April 1998;  1 male(35.4 × 32.4 mm) ( ZRC 2000.2263), Iriomote Island, Ryukyu Islands, coll. Y. Cai& N.K. Ng,  15 June 2000.  TAIWAN– 1 male(28.3 × 25.7 mm) ( ZRC 2001.0025), near Tashi, northern Taiwan, coll. K.-X. Lee, 2000;  1 male(26.2 × 23.9 mm) ( ZRC 2009.0871), Meilunstream, Hualien, coll. H.-C. Liu,  27 January 2002;  1 male, 1 female( ZRC 2000.1847), Hsinfengmangroves, Hsinchu, coll. H.-C. Liu& C.D. Schubart,  17 September 1999;  1 male( ZRC 2002.0414), Lion Museum, Peikuan, Toucheng townarea, coll. H.-C. Liu,  21 June 2002.  CHINA– 2 males( ZRC 2010.0326), purchased, aquarium trade, supposedly from Guangzhou, China, coll. P.K.L. Ng, 2010.  HONG KONG– 2 males(29.6 × 27.6 mm, 30.0 × 26.8 mm) ( ZRC 2002.0223), coll. H.H. Tan,  July 2000;  2 males(24.7 × 21.5 mm, 22.5 × 19.5 mm), 1 female(24.8 × 21.2 mm) ( ZRC1975.6.30.12–14), Tai Po, New Territories, coll. C.L. Soh,  8 June 1975;  6 males, 2 females( ZRC 2019.0539), back mangroves, ca. 22.492306°N 114.036169°E, Mai Po Nature Reserve, coll. S. Cannici, K. Wong, et al.  24 May 2019.   Diagnosis. Carapace distinctly quadrate to transversely rectangular; lateral margin crenulated to entire but no trace of epibranchial tooth; posterolateral margins subparallel or gently converging towards posterior carapace margin; dorsal surface relatively lower, gastric region less swollen, grooves separating regions deep, prominent; frontal lobes separated by deep, broad concavity; ambulatory merus relatively broader; chitinous part of G1 with basal part slightly constricted, distal part relatively broader. In life, carapace dark grey to brown, often with dark median patches, chelae grey to white, with proximal parts sometimes purplish.  Colour. The dorsal carapace surface of fresh  O. dehaaniis grey to dark purplish-brown, often with black or dark blotches on the gastric and adjacent areas ( Fig. 53A, C). The chelipeds vary from white to pale purple (see also Miyake, 1983; Minemizu, 2000; Liu & Wang, 2010; Li & Chiu, 2013; Toyota & Seki, 2014).   Remarks. The type series is represented by 11 specimens( Fransen et al., 1997), and the lectotypewas selected by  Yamaguchi& Baba (1993). This well-known species has a wide range from Japanto southern China. The distribution is interesting as it seems to bracket the range of  O. neglectumfrom Shanghaiand Taiwan. The present specimens from Hong Kongare all easily referrable to  O. dehaanias presently defined, confirming the reports of Soh (1978). The record of  S. hanseni Rathbun, 1897, from the West Indies is clearly incorrect ( Abele, 1975: 48, figs. 1, 2), and from her description, it seems clear that this species is conspecific with  S. dehaani.   Biology.  Orisarma dehaaniis typically associated with coastal rocky supralittoral habitats, living under rocks, or more often in deep burrows in the soil. They may be present in habitats over 100 metres from the shore or along coastal streams. This contrasts with the closely related  O. neglectumthat appears to prefer muddier and more open littoral habitats.  Naraet al. (2006)discussed the social behaviour in this species in Japan, with Hong et al. (2010)commenting on its ecology from Korea. The larval development has been described by Baba & Miyata (1971). Nematode parasites have been reported on by Yoshimura (1990).   Distribution. Koreato Japan, Ryukyus, Taiwan, southern part of mainland China. 3763237534 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC v. Siebold Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237327 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC v. Siebold Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237439 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC J. Takayama Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237385 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC T. Komai Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237418 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC T. Sakai Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237481 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC T. Sakai Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237332 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC Y. Matsuzawa Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237380 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC H. Suzuki Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237502 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC P.K.L. Ng et al. Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237320 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC T. Komai Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237471 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC T. Naruse et al. Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237441 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC P.K.L. Ng et al. Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237489 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC N. Shikatani Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237445 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC Y. Cai & N.K. Ng Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237429 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC K.-X. Lee Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237335 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC H.-C. Liu Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237405 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC H.-C. Liu & C.D. Schubart Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237511 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC H.-C. Liu Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237351 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC P.K.L. Ng Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237389 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC H.H. Tan Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237509 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC C.L. Soh Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype 3763237484 1975-06-08 2019-05-24 1975-06-08 RMNH, ZRC S. Cannici, K. Wong, et al. Japan Japan 22.492306 Kisarazu 1 114.03617 Obitsu River 15 906 D 157, CBM-ZC 165, CBM-ZC 169, CBM-ZC 11406, CBM-ZC 11458, CBM-ZC 7576, ZRC 2002.0226, ZRC 2012.0057, CBM-ZC 3129, ZRC 2011.1026, ZRC 2011.1027, CBM-ZC 9870, ZRC 2000.2263, ZRC 2001.0025, ZRC 2009.0871, ZRC 2000.1847, ZRC 2002.0414, ZRC 2010.0326, ZRC 2002.0223, ZRC 2019.0539 48 12 36 Chiba lectotype