Sesarma johorensis Tweedie, 1940: 103 Tweedie, 1950: 342 Sesarma ( Sesarma ) johorensis Serène, 1968: 105 Pseudosesarma johorense Tan & Ng, 1994: 82 “ Pseudosesarma ” johorense Ng et al., 2008a: 220 Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène & Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species Schubart, Christoph D. Ng, Peter K. L. Raffles Bulletin of Zoology 2020 2020-12-23 68 891 994 84GYW (Tweedie, 1940) Tweedie 1940 [924,1340,413,437] Malacostraca Sesarmidae Manarma Animalia Decapoda 73 964 Arthropoda species johorense  ( Figs. 24D, 26B, 29C, 33B, E, 41F–J)      Sesarma johorensis Tweedie, 1940: 103, fig. 9, pl. 24-3;  Tweedie, 1950: 342.    Sesarma( Sesarma) johorensis–  Serène, 1968: 105.    Pseudosesarma johorense–  Tan & Ng, 1994: 82.   “ Pseudosesarma” johorense–  Ng et al., 2008a: 220.   Material examined.  Paratypes: male (10.0 × 8.5 mm), 1 female(9.6 × 8.3 mm) ( ZRC1965.7.29.189–190), Pendas River, southern Johor, Peninsular Malaysia, coll. M.W.F. Tweedie,  February 1937;  1 female(11.2 × 9.1 mm) ( ZRC1970.3.13.2), Pendas River, southern Johor, Peninsular Malaysia, coll. M.W.F. Tweedie,  February 1937.  Others: SINGAPORE– 1 male(14.6 × 12.6 mm) ( ZRC1971.9.24.14), Sungei Seletar, coll. C.L. Soh,  31 December 1966;  1 female( ZRC1971.9.24.13), Simpang River, Mak Wai, Singapore, coll. C.L. Soh,  3 March 1966;  1 female( ZRC1971.9.22.8), Simpang River, Mak Wai, coll. C.L. Soh,  20 December 1966;  1 female( ZRC1973.11.2.496), Sungei Seletar, coll.  14 June 1967;  3 males( ZRC1971.9.24.12), Sungei Seletar, coll. C.L. Soh,  31 December 1966;  1 juvenile( ZRC1970.2.23.4), Sungei Seletar, coll. C.L. Soh,  31 December 1966;  1 female( ZRC1970.2.23.1), Sungei Seletar, coll. C.L. Soh,  11 December 1966;  1 male( ZRC1971.9.22.9), Sungei Seletar, coll. C.L. Soh,  31 December 1966;  1 female( ZRC1971.9.24.11), Sungei Seletar, coll. C.L. Soh,  31 December 1966. PENINSULAR MALAYSIA– 1 female( ZRC1965.7.29.57), Sedili River, Johor,  March 1938.  LABUAN– 4 males( ZRC1965.7.29.185–188), coll. M.W.F. Tweedie, 1938.  VIETNAM– 1 male( ZRC1969.10.1.4), Cantho, coll. Nguyen Thi Lai,  August 1969;  2 juveniles( ZRC1970.2.23.2–3), Cantho, coll. Institute of Oceanography, Nhatrang, 1958.   Diagnosis. Dorsal surface of carapace relatively flatter, regions relatively less prominent; fingers of cheliped relatively shorter in adult males; male pleonal somite 6 relatively longer, telson more rounded; distal chitinous part of G1 gently curved distally. In life, carapace uniform dark brown.  Colour. According to Tweedie (1940: 105), the “colour of the carapace is very dark greenish brown and of the chelae, pale yellow”.   Remarks. The differences between  M. johorensisand  M. moeschiihave been discussed under the latter species.   Biology. The species lives in mangrove swamps near the main river ( Tweedie, 1940: 103).   Distribution. Known only from southern Peninsular Malaysia, Sarawak, and Vietnam( Tweedie, 1940, 1950; present data). 3763237304 1937-02 ZRC M. W. F. Tweedie Peninsular Malaysia Pendas River southern Johor 73 964 1 1 paratype 3763237564 1937-02 ZRC M. W. F. Tweedie Peninsular Malaysia Pendas River southern Johor 73 964 1 1 paratype 3763237374 1966-12-31 ZRC C. L. Soh Singapore Others 73 964 1 1 paratype 3763237355 1966-03-03 ZRC C. L. Soh Singapore Singapore Simpang River Mak Wai 73 964 1 1 paratype 3763237517 1966-12-20 ZRC C. L. Soh Singapore Simpang River Mak Wai 73 964 1 1 paratype 3763237533 1967-06-14 ZRC Singapore Sungei Seletar 73 964 1 1 paratype 3763237403 1966-12-31 ZRC C. L. Soh Singapore Sungei Seletar 73 964 3 3 paratype 3763237350 1966-12-31 ZRC C. L. Soh Singapore Sungei Seletar 73 964 1 1 paratype 3763237396 1966-12-11 ZRC C. L. Soh Singapore Sungei Seletar 73 964 1 1 paratype 3763237338 1966-12-31 ZRC C. L. Soh Singapore Sungei Seletar 73 964 1 1 paratype 3763237528 1966-12-31 ZRC C. L. Soh Singapore Sungei Seletar 73 964 1 1 paratype 3763237440 1938-03 ZRC Singapore Sedili River 73 964 1 1 Johor paratype 3763237547 ZRC M. W. F. Tweedie Vietnam LABUAN 73 964 4 4 Labuan paratype 3763237553 1969-08 ZRC M. W. F. Tweedie Vietnam LABUAN 73 964 4 4 Labuan paratype 3763237544 ZRC Nhatrang Vietnam Cantho 73 964 2 2 Labuan paratype