Brocchi 1814 : 289 Hoernes 1878a : 195 Hoernes & Auinger 1879 : 49 Conus ( Chelyconus ) vindobonensis var. mariae Kojumdgieva & Strachimirov 1960 : 213 Conus ( Chelyconus ) mariae Hoernes et Auinger, 1879 Hinculov 1968 : 150 Conus vindobonensis mariae (Hoernes et Auinger) Ionesi & Nicorici 1994 : 62 Varioconus pelagicus ( Brocchi, 1814 ) Landau et al . 2013 : 247 Varioconus pelagicus ( Brocchi, 1814 ) Kovács & Vicián 2013 : 87 Conus pelagicus Brocc. Lautoconus ventricosus ( Gmelin, 1791 ) Lautoconus pelagicus Conus mediterraneus Hwass in Bruguière, 1792 Lautoconus ventricosus ( Gmelin, 1791 ) Conus ( Chelyconus ) pelagicus incertula Conus incertula Sacco, 1893b Lautoconus ventricosus Plagioconus Lautoconus pelagicus Conus suessi Hoernes & Auinger (2013) L. pelagicus Plagioconus Leporiconus suessi ( Hoernes & Auinger, 1879 ) A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea Harzhauser, Mathias Landau, Bernard Zootaxa 2016 4210 1 1 178 Brocchi, 1814 Brocchi 1814 [151,612,1196,1222] Gastropoda Conidae Lautoconus CoL Animalia Neogastropoda 83 84 Mollusca species pelagicus     Conus pelagicusnob.—  Brocchi 1814: 289, pl. 2, fig. 9.  [  Chelyconus] [  Conus]  Mariaen. f.—  Hoernes 1878a: 195(nomen nudum).  C.[ onus]  Mariae— Hoernes 1878b: 207 (nomen nudum).    Conus( Chelyconus) Mariaenov. form.—  Hoernes & Auinger 1879: 49, pl. 6, fig. 7.     Conus( Chelyconus) vindobonensisvar. mariae( Hoernes und Auinger 1879)—Kojumdgieva in  Kojumdgieva & Strachimirov 1960: 213, pl. 50, fig. 6.    Conus( Chelyconus) mariae Hoernes et Auinger, 1879—  Hinculov 1968: 150, pl. 37, figs 17a–b.    Conus vindobonensis mariae(Hoernes et Auinger)—  Ionesi & Nicorici 1994: 62, pl. 5, fig. 8.    Varioconus pelagicus( Brocchi, 1814)—  Landau et al. 2013: 247, pl. 39, figs 6–7, pl. 41, fig. 14, pl. 42, fig. 8, pl. 82, figs 1–2.    Varioconus pelagicus( Brocchi, 1814)—  Kovács & Vicián 2013: 87, figs 131–137. non  Conus pelagicusBrocc.—Hörnes 1851: 31, pl. 3, figs 4a–d [=  Lautoconus ventricosus( Gmelin, 1791)].     Typematerial. Lectotype(following ICZNArticle 74.6) stored in the Museo Civico di Storia Naturale di Milano, Catalogue number 4668, Piacentino, Italy, Pliocene; illustrated in Pinna & Spezia (1978; pl. 18, fig. 4). The type locality of  Conus mariae Hoernes & Auinger, 1879, which is considered to be a junior synonym of  C. pelagicus, is the middle Miocene locality Bujtur in Romania.   Studied material.1 spec.  NHMW1861/0035/0004, Bujtur( Romania), illustrated in  Hoernes& Auinger(1879, pl. 6, fig. 7); 2 spec.   NHMW1868/0001/0387, Lăpugiu de Sus( Romania).    Illustratedmaterial.  Figs19A 1–A4, 17J: Bujtur( Romania): SL: 53.6 mm, MD: 24.9 mm,  NHMW1861/0035/ 0 0 0 4, illustrated in Hoernes & Auinger (1879, pl. 6, fig. 7).   FIGURE 19A1–A4.  Lautoconus pelagicus(Brocchi, 1814), Bujtur (Romania), NHMW 1861/0035/0004. 19B1–B3.  Lautoconus ponderosus(Brocchi, 1814), Steinebrunn (Austria), NHMW 1846/0037/0064. 19C1–C3.  Lautoconus ponderosus(Brocchi, 1814), Steinebrunn (Austria), GBA 1856/004/0001/01.  Revised description.Medium-sized, biconical, solid, glossy shells with high conical spire; early spire whorls probably tuberculate; spire whorls flat to weakly convex, not striate; suture narrow but deeply impressed, almost canaliculated. Subsutural flexure shallow, moderately curved, moderately asymmetrical; last spire whorl weakly concave; indistinct angulation on shoulder; last whorl slightly ventricose, constricted at base. No spiral sculpture except for several spiral cords on base. Aperture moderately narrow. Siphonal fasciole strongly swollen, convex, twisted; inner lip narrow, straight; siphonal canal moderately long, reflected. Colour pattern under UV light consisting of flammulae on shoulder and spire and irregularly spaced short dashes on the entire last whorl, forming three broad bands separated by two fluorescing bands.   Shell measurements and ratios. Three specimens are available: largest specimen SL: 57.6 mm, MD: 28.2 mm, spire angle: 82/85/83°, last whorl angle: 33/35/32, LW: 2.04/2.09/2.15, RD: 0.59/0.59/0.56, PMD: 0.86/0.88/ 0.91, RSH: 0.17/0.18/0.17; spire angle: 95°, last whorl angle: 37°; second specimen: SL: 50.4 mm, MD: 25.2 mm, LW: 2.0, RD: 0.64, PMD: 0.85, RSH: 0.22.   Discussion.Differs from the somewhat similar  Conus s.l.  posticestriatus(Kojumdgieva in Kojumdgieva & Strachimirov 1960)by the absence of any spiral grooves on the spire whorls, the constricted base and the reflected siphonal canal. Hall (1894) and Landau et al. (2013)synonymized  Conus mariae Hoernes & Auinger, 1879with  Lautoconus pelagicus( Brocchi, 1814), which is followed herein.    Lautoconus pelagicuswas also reported from the ViennaBasin by Hörnes (1851, pl. 3, fig. 4). Later, Hoernes & Auinger (1879)recognized this specimen as a recent  Conus mediterraneusHwass in Bruguière, 1792[=  Lautoconus ventricosus( Gmelin, 1791)]. Although Sacco (1893b)was aware of that statement by Hoernes & Auinger (1879), he doubted that the question was solved and introduced  Conus( Chelyconus) pelagicus incertulaas new variation name. The specimen is still stored in the GBA collection (GBA 1856/0004/0002). Preservation and colour pattern reveal this specimen as a Recent shell, as supposed by Hoernes & Auinger (1879); moreover, the preservation mode differs completely from the fossil shells from Baden, which is given as the locality on the label. Consequently,  Conus incertula Sacco, 1893bis asubjective junior synonym of  Lautoconus ventricosus. The species was discussed again by Hall (1966)by synonymising also  Plagioconuslapugyensis( Hoernes & Auinger, 1879) with  Lautoconus pelagicus. In addition, Kovács & Vicián (2013)and Landau et al. (2013)treated  Conus suessiHoernes & Auinger (2013)as a junior synonym of  L. pelagicus. Both synonymizations are rejected herein [see discussions on  Plagioconuslapugyensis( Hoernes & Auinger, 1879) and  Leporiconus suessi( Hoernes & Auinger, 1879)].   Paleoenvironment.The occurrences in the Turkish Karman Basin suggest shallow marine environments.   Distribution in Paratethys.Badenian (middle Miocene):  ViennaBasin:Gainfarn ( Austria), Mikulov- Kienberk, Hrušovany ( Czech Republic) ( Hoernes & Auinger 1879); Eisenstadt-Sopron Basin:Forchtenau ( Austria) ( Sieber 1956); Pannonian Basin:Letkés ( Hungary) ( Kovács & Vicián 2013); Transylvanian Basin: Lăpugiu de Sus, Bujtur ( Romania) ( Hoernes & Auinger 1879);  BuzăuBasin: Valea Muscel ( Romania) ( Ionesi & Nicorici 1994); Caransebeş-Mehadia Basin:Calvei ( Romania) ( Hinculov 1968); Dacian Basin:Opanec ( Bulgaria) ( Kojumdgieva & Strachimirov 1960).   Proto-Mediterranean Sea and north eastern Atlantic.Burdigalian (early Miocene): Turin Hills ( Italy); Langhian (middle Miocene): AquitaineBasin ( France); Serravallian (middle Miocene): KaramanBasin ( Turkey); Tortonian (late Miocene): Po Basin (Italy); early Pliocene: western Mediterranean, Estepona Basin ( Spain), central Mediterranean( Italy); late Pliocene: central Mediterranean( Italy) (see Landau et al. 2013for details). 1425219515 ICZN Museo Civico di Storia Naturale di Milano Italy Italy 83 84 2 lectotype 1425219502 NHMW Bujtur & Hoernes & Auinger Romania Romania 83 84 NHMW 1861 1 1425219493 [462,1063,1889,1914] NHMW Lapugiu de Sus Romania Romania 83 84 NHMW 1868 1 1425219510 NHMW Romania Bujtur Illustrated Figs 83 84 NHMW 1861 1 1425219496 [199,1431,727,752] Italy north eastern Atlantic. Burdigalian 85 86 1 1425219499 [151,773,764,789] France Aquitaine 85 86 1 Aquitaine 1425219504 [785,1432,764,789] Turkey Karaman 85 86 1 Karaman 1425219514 Spain western Mediterranean central Mediterranean 85 86 1 1425219534 [406,1293,836,861] Italy central Mediterranean 85 86 1