A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea Harzhauser, Mathias Landau, Bernard Zootaxa 2016 4210 1 1 178 XK29 [151,454,1891,1917] Gastropoda Conidae Conasprella Animalia Neogastropoda 41 42 Mollusca species minutissima     Holotype: Figs  4I1–I5: SL: 13.1, MD: 6.3 mm,  NHMW1869/0001/0331a, Steinebrunn( Austria).    Paratype: Figs 4H1–H3: SL: 10.6, MD: 5.1 mm,  NHMW1869/0001/0331b, Steinebrunn( Austria).     Paratype: Figs 3E, 4G1–G3: SL: 10.4, MD: 4.7 mm,  NHMW1869/0001/0331c, Steinebrunn( Austria).   Additional material:2 spec. NHMW 1869/0001/0331d, 1 spec.   NHMW1858/0029/0017, Steinebrunn( Austria); 1 spec.  NHMW1870/0054/0044, Niederleis( Austria); 1 spec.  NHMW1869/0001/0627, Marz( Austria).  Type stratum:Bioclastic sand and marl of the Baden Group.    Typelocality: Steinebrunn( Austria).  Age:Middle Miocene, middle Badenian (= late Langhian).   Etymology.Referring to the small size.   Description.Very small, slender biconical shells. High, scalariform spire of at least 7 teleoconch whorls. Spire whorls carinate just above suture, weakly tuberculate; sutural ramp distinctly striate. Impressed, somewhat irregular suture. Last whorl with flat to faintly concave sutural ramp, weakly striate. Subsutural flexure shallow, strongly curved, moderately asymmetrical. Last whorl elongate conical to weakly ventricose below angulated shoulder; slightly constricted at base. Aperture with subparallel margins; short wide siphonal canal. Siphonal fasciole obsolete. Sculpture consisting of about 15–20 spiral cords with spirally elongate beads. Spiral sculpture overlays much weaker growth lines forming a faintly cancellate pattern. Spirals become crowded on base where beads are only weakly developed. Colour pattern in UV light consisting of a few large, widely-spaced subquadratic blotches on sutural ramp of last whorl and weaker blotches on spire whorls; beads on last whorl appear as dots on thin spirals.  Shell measurements and ratios.n = 5: largest specimen: SL: 13.1 mm, MD: 6.3 mm, mean SL: 10.4 (σ = 0.8), mean MD: 5.1 (σ = 0.7), spire angle: µ = 63.2° (σ = 8.5°), last whorl angle: µ = 32.4° (σ = 2.4°), LW: µ = 2.04 (σ = 0.13), RD: µ = 0.68 (σ = 0.03), PMD: µ = 0.81 (σ = 0.09), RSH: µ = 0.28 (σ = 0.04).   Discussion.This tiny species was mixed with  Artemidiconus stacheiin the NHMW collections, probably due to its small size and beaded sculpture. The scalariform spire and the lack of spiral cords on the spire whorls, however, allow a clear separation.  Conasprella berwerthi( Hoernes & Auinger, 1879)develops a similar but coarser sculpture on the last whorl. Moreover, it differs in its lower spire, the much broader last whorl and it lacks subquadratic blotches on the sutural ramp.  Phasmoconus ottiliae( Hoernes & Auinger, 1879)lacks the scalariform spire outline, has an elongate last whorl, a denser spiral sculpture, lacks the prominent beads and differs in its colour pattern of dense spirals.  Paleoenvironment.The occurrence at Steinebrunn suggests seagrass meadows in shallow marine nearshore settings as habitat.   Distribution in Paratethys.Badenian (middle Miocene):  ViennaBasin:Niederleis ( Austria), Steinebrunn ( Austria); Eisenstadt SopronBasin: Marz, Forchtenau ( Austria). 1425219447 [151,1237,2000,2029] 4 I NHMW Austria Steinebrunn 41 42 NHMW 1869 1 holotype 1425219460 [199,1309,151,180] NHMW Austria Steinebrunn 42 43 NHMW 1869 1 paratype 1425219461 [199,1352,190,219] NHMW Austria Steinebrunn 42 43 NHMW 1869 1 paratype 1425219455 NHMW Austria Steinebrunn 42 43 NHMW 1858 1 1425219459 [266,872,265,290] NHMW Austria Niederleis 42 43 NHMW 1870 1 1425219471 [883,1433,265,290] NHMW Austria Marz 42 43 NHMW 1869 1 1425219457 [199,618,337,362] Austria Steinebrunn 42 43 1 holotype